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red spotted clarkia, red spotted or redspot clarkia, redspot clarkia

arewell-to-spring, farewell-to-spring, farewell-to-spring clarkia, godetia, yellow clarkia

Stems

erect or decumbent, to 60 cm, puberulent.

erect to decumbent, 20–200 cm, puberulent.

Leaves

petiole 0–5 mm;

blade linear to lanceolate, 1–6 cm.

petiole to 10 mm;

blade linear to lanceolate, 1–6 cm.

Inflorescences

open or dense racemes or panicles, axis straight;

buds erect.

open or dense spikes or racemes, axis straight;

buds erect.

Flowers

floral tube 5–15 mm;

sepals usually reflexed individually or in pairs, or rarely together to one side, tips not distinct in bud;

corolla bowl-shaped, petals lavender to wine-red, often shading white or pale yellow toward base, unspotted or with bright red or purplish red spot near or proximal to middle, 10–25 mm;

stamens 8, subequal;

ovary 8-grooved;

stigma exserted beyond anthers.

floral tube 3–10 mm;

sepals usually reflexed together to one side, or rarely in pairs or singly;

corolla bowl-shaped, petals pale pink to lavender, usually with red spot or mark near middle, obovate to fan-shaped, 15–60 mm, not lobed, apex sometimes shallowly notched or erose;

stamens 8, in 2 subequal sets;

ovary cylindrical and 4-grooved or fusiform and 8-grooved, puberulent;

stigma exserted or not beyond anthers.

Capsules

15–20 mm.

15–40 mm, sometimes broader distally;

pedicel 0–13 mm.

Seeds

brown or gray, 0.7–1 mm, scaly, crest 0.1 mm.

brown to grayish brown, 1–1.5 mm, scaly, crest 0.1 mm.

Clarkia speciosa

Clarkia amoena

Distribution
from USDA
California
[WildflowerSearch map]
[BONAP county map]
from USDA
w North America
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 4 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 5 (5 in the flora).

Clarkia amoena is closely related to C. rubicunda, which differs morphologically mainly in the color pattern of the petals. Petals of C. amoena have a conspicuous red spot or group of small red spots or marks near the middle, whereas those of C. rubicunda have a red area at the base and are not spotted near the middle. The areas of distribution of the two species barely overlap in California just north of San Francisco Bay, and C. rubicunda could be considered a southern geographical race or subspecies of C. amoena were it not that their readily formed hybrids are sterile due to chromosomal rearrangement. Clarkia amoena is one of the parent species of the allotetraploid C. gracilis. Intermediates between subspecies are frequent.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals without red or purplish red spot.
→ 2
2. Stems erect to decumbent; petals dark purplish red to lavender, ± white or pale yellow near base, rarely pale yellow throughout.
subsp. speciosa
2. Stems decumbent; petals lavender to pink shading pale yellow or white in proximal 1/2.
subsp. immaculata
1. Petals with bright red or purplish red spot near or proximal to middle.
→ 3
3. Inflorescences dense racemes.
subsp. nitens
3. Inflorescences open racemes or panicles.
→ 4
4. Petals usually with bright red spot; branches on well-developed plants many, few-flowered.
subsp. speciosa
4. Petals with purplish red spot; branches on well-developed plants virgate, few, many- flowered.
subsp. polyantha
1. Stigmas not exserted beyond anthers; petals less than 20 mm; coastal British Columbia, Oregon, Washington.
subsp. caurina
1. Stigmas exserted beyond anthers; petals 15–60 mm.
→ 2
2. Ovaries 8-grooved, broadly fusiform, 8–12 mm wide.
subsp. whitneyi
2. Ovaries 4-grooved, cylindrical to subclavate, 2–6 mm wide.
→ 3
3. Inflorescences open, internodes longer than subtending flowers, bracts sublinear.
subsp. huntiana
3. Inflorescences congested, internodes shorter than subtending flowers, bracts narrowly lanceolate or wider.
→ 4
4. Stems decumbent to suberect, to 100 cm; petals 20–35 mm, usually with red spot mid blade; coastal California.
subsp. amoena
4. Stems erect, to 200 cm; petals 30–40 mm, usually without red spot or with very small spot or streak mid blade; inland Oregon, Washington.
subsp. lindleyi
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Godetia Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Rhodanthos > subsect. Primigenia
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. speciosa subsp. immaculata, C. speciosa subsp. nitens, C. speciosa subsp. polyantha, C. speciosa subsp. speciosa
C. amoena subsp. amoena, C. amoena subsp. caurina, C. amoena subsp. huntiana, C. amoena subsp. lindleyi, C. amoena subsp. whitneyi
Synonyms Oenothera viminea var. parviflora, Godetia parviflora Oenothera amoena, Godetia amoena
Name authority H. Lewis & M. E. Lewis: Madroño 12: 34. (1953) (Lehmann) A. Nelson & J. F. Macbride: Bot. Gaz. 65: 62. (1918)
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