Clarkia sect. Eucharidium
Onagraceae subfam. onagroideae
stipules present or absent.
axis suberect or slightly recurved;
buds pendent.
floral tube narrowly tubular, 13–35 mm;
sepals reflexed together to 1 side;
petals pink, sometimes white-streaked, fan-shaped, conspicuously 3-lobed, middle lobe often longer than laterals, claw slender, not lobed;
stamens 4.
floral tube present or, rarely, absent;
sepals 2 or 4 (very rarely 3), deciduous with floral tube, petals, and stamens;
petals yellow, white, pink, red, rarely in combination.
subterete;
sessile or subsessile.
Clarkia sect. Eucharidium
Onagraceae subfam. onagroideae
Species 2 (2 in the flora).
Section Eucharidium includes two species characterized by large tri-lobed petals, four rather than eight stamens, and a long floral tube that adapts them to pollination by long-tongued Lepidoptera or Diptera (G. A. Allen et al. 1990).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera 21, species 582 (16 genera, 246 species in the flora).
Onagroideae encompass the main lineage of the family, after the early branching of Ludwigia (R. A. Levin et al. 2003, 2004). This large and diverse lineage is distinguished by the presence of a floral tube beyond the apex of the ovary; sepals deciduous with the floral tube, petals, and stamens; pollen shed in monads (or tetrads in Chylismia sect. Lignothera and all but one species of Epilobium); ovular vascular system exclusively transseptal (R. H. Eyde 1981); ovule archesporium multicellular (H. Tobe and P. H. Raven 1996); and change in base chromosome number from x = 8 in Ludwigia to x = 10 or x = 11 at the base of Onagroideae (Raven 1979; Levin et al. 2003). Molecular work (Levin et al. 2003, 2004) substantially supports the traditional tribal classification (P. A. Munz 1965; Raven 1979, 1988); tribes are recognized to delimit major branches within the phylogeny of Onagroideae, where the branches comprise strongly supported monophyletic groups of one or more genera.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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