Clarkia rubicunda
Onagraceae tribe Onagreae
farewell-to-spring, ruby chalice clarkia
erect or decumbent, to 150 cm, puberulent;
buds erect.
petiole to 10 mm;
blade lanceolate to elliptic, 1–4 cm.
alternate or basal;
stipules absent.
open or dense spikes or racemes, axis straight;
buds erect.
floral tube 4–10 mm;
sepals reflexed together to one side;
corolla bowl-shaped, petals pink to lavender, base red or purplish red, fan-shaped, 10–30 mm, apex erose;
stamens 8, subequal;
ovary cylindrical, 4-grooved, puberulent;
stigma exserted beyond anthers.
usually actinomorphic, rarely slightly zygomorphic (in Oenothera), (3 or)4-merous;
stamens 2 times as many, or rarely as many, as sepals;
pollen usually shed in monads, rarely tetrads (Chylismia sect. Lignothera).
a dry capsule, usually dehiscent, sometimes indehiscent.
20–40 mm;
pedicel 0–25(–40) mm.
brown to grayish brown, 1.2–1.5 mm, scaly, crest 0.2 mm.
few to numerous, without hairs or wings, [very rarely with asymmetrical dry wing (Xylonagra)], or with dry (Oenothera), erose or smooth wing, or with thick, papillate wings (Chylismiella).
= 14.
Clarkia rubicunda
Onagraceae tribe Onagreae
Clarkia rubicunda is known from the central coast of California, from Contra Costa and Marin counties south along the coast and foothills to northern San Luis Obispo County.
Clarkia rubicunda is probably a derivative of C. amoena and may be ancestral to C. franciscana. Clarkia rubicunda is distinguishable from some populations of C. amoena only by the absence of a red spot or group of spots near the middle of the petal and the presence of a red area at the base of the petal. Clarkia rubicunda can be distinguished from C. franciscana by the position of the stigma and size and shape of the petals. All three species differ in chromosome arrangement and hybrids are highly sterile.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera 13, species 265 (12 genera, 199 species in the flora).
Onagreae account for more than half the total genera in Onagraceae and diversified from a center in southwestern North America (L. Katinas et al. 2004). Delimitation of the tribe by W. L. Wagner et al. (2007) differs from previous ones by the exclusion of Gongylocarpus, now in its own tribe, by the segregation of eight genera (Camissoniopsis, Chylismia, Chylismiella, Eremothera, Eulobus, Neoholmgrenia, Taraxia, and Tetrapteron) from Camissonia, and by the inclusion of three previously separate genera (Calylophus, Gaura, and Stenosiphon) in Oenothera. Within the branch of the family that lacks stipules (Gongylocarpeae, Epilobieae, and Onagreae), the last two tribes form a clade that has very strong molecular support (R. A. Levin et al. 2003, 2004), but no obvious morphological synapomorphy. The clade may be defined by a cytogenetic change from the base chromosome number of x = 11 found in Circaeeae, Gongylocarpeae, and Lopezieae, to x = 18 in Epilobieae, and x = 7 in Onagreae; however, these changes could also have occurred independently. Other than the new chromosome number x = 7, the only apparent morphological synapomorphy for Onagreae alone is pollen with prominent apertural protrusions (J. Praglowski et al. 1987, 1989), a character state also found in Circaeeae (Praglowski et al. 1994). The monophyly of Onagreae has moderate (Levin et al. 2004) to strong support (V. S. Ford and L. D. Gottlieb 2007).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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