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farewell-to-spring, ruby chalice clarkia

arewell-to-spring, farewell-to-spring, farewell-to-spring clarkia, godetia, yellow clarkia

Stems

erect or decumbent, to 150 cm, puberulent;

buds erect.

erect to decumbent, 20–200 cm, puberulent.

Leaves

petiole to 10 mm;

blade lanceolate to elliptic, 1–4 cm.

petiole to 10 mm;

blade linear to lanceolate, 1–6 cm.

Inflorescences

open or dense spikes or racemes, axis straight;

buds erect.

open or dense spikes or racemes, axis straight;

buds erect.

Flowers

floral tube 4–10 mm;

sepals reflexed together to one side;

corolla bowl-shaped, petals pink to lavender, base red or purplish red, fan-shaped, 10–30 mm, apex erose;

stamens 8, subequal;

ovary cylindrical, 4-grooved, puberulent;

stigma exserted beyond anthers.

floral tube 3–10 mm;

sepals usually reflexed together to one side, or rarely in pairs or singly;

corolla bowl-shaped, petals pale pink to lavender, usually with red spot or mark near middle, obovate to fan-shaped, 15–60 mm, not lobed, apex sometimes shallowly notched or erose;

stamens 8, in 2 subequal sets;

ovary cylindrical and 4-grooved or fusiform and 8-grooved, puberulent;

stigma exserted or not beyond anthers.

Capsules

20–40 mm;

pedicel 0–25(–40) mm.

15–40 mm, sometimes broader distally;

pedicel 0–13 mm.

Seeds

brown to grayish brown, 1.2–1.5 mm, scaly, crest 0.2 mm.

brown to grayish brown, 1–1.5 mm, scaly, crest 0.1 mm.

2n

= 14.

Clarkia rubicunda

Clarkia amoena

Phenology Flowering May–Aug.
Habitat Openings in woodlands, forests, chaparral, coastal scrub.
Elevation 0–500 m. (0–1600 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from USDA
w North America
[WildflowerSearch map]
[BONAP county map]
Discussion

Clarkia rubicunda is known from the central coast of California, from Contra Costa and Marin counties south along the coast and foothills to northern San Luis Obispo County.

Clarkia rubicunda is probably a derivative of C. amoena and may be ancestral to C. franciscana. Clarkia rubicunda is distinguishable from some populations of C. amoena only by the absence of a red spot or group of spots near the middle of the petal and the presence of a red area at the base of the petal. Clarkia rubicunda can be distinguished from C. franciscana by the position of the stigma and size and shape of the petals. All three species differ in chromosome arrangement and hybrids are highly sterile.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 5 (5 in the flora).

Clarkia amoena is closely related to C. rubicunda, which differs morphologically mainly in the color pattern of the petals. Petals of C. amoena have a conspicuous red spot or group of small red spots or marks near the middle, whereas those of C. rubicunda have a red area at the base and are not spotted near the middle. The areas of distribution of the two species barely overlap in California just north of San Francisco Bay, and C. rubicunda could be considered a southern geographical race or subspecies of C. amoena were it not that their readily formed hybrids are sterile due to chromosomal rearrangement. Clarkia amoena is one of the parent species of the allotetraploid C. gracilis. Intermediates between subspecies are frequent.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stigmas not exserted beyond anthers; petals less than 20 mm; coastal British Columbia, Oregon, Washington.
subsp. caurina
1. Stigmas exserted beyond anthers; petals 15–60 mm.
→ 2
2. Ovaries 8-grooved, broadly fusiform, 8–12 mm wide.
subsp. whitneyi
2. Ovaries 4-grooved, cylindrical to subclavate, 2–6 mm wide.
→ 3
3. Inflorescences open, internodes longer than subtending flowers, bracts sublinear.
subsp. huntiana
3. Inflorescences congested, internodes shorter than subtending flowers, bracts narrowly lanceolate or wider.
→ 4
4. Stems decumbent to suberect, to 100 cm; petals 20–35 mm, usually with red spot mid blade; coastal California.
subsp. amoena
4. Stems erect, to 200 cm; petals 30–40 mm, usually without red spot or with very small spot or streak mid blade; inland Oregon, Washington.
subsp. lindleyi
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Rhodanthos > subsect. Primigenia Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Rhodanthos > subsect. Primigenia
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. amoena subsp. amoena, C. amoena subsp. caurina, C. amoena subsp. huntiana, C. amoena subsp. lindleyi, C. amoena subsp. whitneyi
Synonyms Godetia rubicunda, C. rubicunda subsp. blasdalei, G. blasdalei Oenothera amoena, Godetia amoena
Name authority (Lindley) H. Lewis & M. E. Lewis: Madroño 12: 34. (1953) (Lehmann) A. Nelson & J. F. Macbride: Bot. Gaz. 65: 62. (1918)
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