Clarkia rhomboidea |
Clarkia affinis |
|
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common clarkia, diamond clarkia, diamond fairyfan, forest clarkia, rhombic petal clarkia, rhomboid farewell-to-spring, tongue clarkia |
chaparral clarkia, chaparral clarkia or fairyfan, chaparral fairyfan |
|
Stems | erect, to 100 cm, puberulent. |
erect, to 80 cm, puberulent. |
Leaves | petiole 5–25 mm; blade lanceolate to elliptic or ovate, 1–6 cm. |
petiole 0–3 mm; blade linear to narrowly lanceolate, 1.5–7 cm. |
Inflorescences | open racemes, axis in bud recurved 1–3 nodes distal to open flowers; buds pendent, narrowly obovoid, tip acute to obtuse, often curved to one side. |
dense spikes, axis straight; buds erect. |
Flowers | floral tube 1–3 mm; sepals reflexed individually; corolla rotate, petals pinkish lavender, often with darker flecks, narrowly to broadly obovate or rhombic, sometimes ± 3-lobed, 6–12(–14) × 3–7 mm; stamens 8, subequal, subtended by ciliate scales, pollen blue-gray; ovary shallowly 4-grooved; stigma not or rarely exserted beyond anthers. |
floral tube1.5–4 mm; sepals reflexed together to 1 side; corolla bowl-shaped, petals 5–15 mm; stamens 8, subequal; ovary cylindrical, 8-grooved, length at least 10 times width; stigma not exserted beyond anthers. |
Capsules | 10–25 mm; pedicel 1–4 mm. |
15–30 mm, beak 3–7 mm; pedicel 0–5 mm. |
Seeds | brown, gray, or mottled, 1–1.5 mm, scaly-echinate, crest 0.1 mm, inconspicuous. |
brown or gray, 1–1.5 mm, scaly, crest 0.1 mm. |
2n | = 24. |
= 52. |
Clarkia rhomboidea |
Clarkia affinis |
|
Phenology | Flowering May–Sep. | Flowering Apr–Jun. |
Habitat | Yellow-pine forests, woodlands. | Openings in woodlands and chaparral. |
Elevation | 0–3000 m. (0–9800 ft.) | 0–500 m. (0–1600 ft.) |
Distribution |
AZ; CA; ID; MT; NV; OR; UT; WA; Mexico (Baja California)
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CA
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Discussion | Clarkia rhomboidea is a tetraploid derived from C. mildrediae and C. virgata or a closely related species. The six diploid species in sect. Myxocarpa closely related to C. rhomboidea (C. australis, C. borealis, C. mildrediae, C. mosquinii, C. stellata, and C. virgata) are California endemics with relatively small areas of distribution, whereas C. rhomboidea occurs throughout much of the western United States and is morphologically much more variable. Clarkia rhomboidea characteristically has relatively small, self-pollinating flowers with the stigma in contact with the anthers. Among the diploid species only C. stellata has similar small, self-pollinating flowers but is distinguished by yellow pollen and petals with a shallowly 3-lobed blade that is not flecked. Rare populations of C. rhomboidea have relatively large flowers with the stigma exserted beyond the anthers. When they occur within the geographical range of the outcrossing diploid species, they may be difficult to distinguish without determining chromosome number. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Clarkia affinis is known primarily from west-central California and the North Coast Ranges, and more scattered in the Sierra Nevada Foothills and Western Transverse Ranges. Clarkia affinis is a hexaploid most closely related to C. purpurea; both have 2n = 52. Chromosome pairing in hybrids between them, as well as morphology, suggest that they have a tetraploid (2n = 34) genome in common. The two species are most readily distinguished by their immature capsules, which in C. affinis are slender, at least ten times longer than wide, beaked, and shallowly grooved, whereas those of C. purpurea are stout, not more than eight times longer than wide, not prominently beaked, and deeply grooved; the sepals of the former are generally reflexed together in fours whereas those of the latter are reflexed individually or in twos. Based on morphology and molecular data, the diploid genome probably came from C. cylindrica or a related species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 10. | FNA vol. 10. |
Parent taxa | ||
Sibling taxa | ||
Name authority | Douglas in W. J. Hooker: Fl. Bor.-Amer. 1: 214. (1832) | H. Lewis & M. E. Lewis: Madroño 12: 34. (1953) |
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