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purple clarkia, winecup clarkia, winecup fairyfan

Vine Hill clarkia

Stems

erect or rarely decumbent, to 100 cm, glabrous and sometimes glaucous or sparsely to densely puberulent, sometimes mixed with longer, spreading hairs.

erect, to 60 cm, glabrous or sparsely puberulent.

Leaves

petiole 0–2 mm;

blade linear or narrowly lanceolate to elliptic or ovate, 1.5–7 cm.

petiole 0–2 mm;

blade lanceolate, 2–2.5 cm.

Inflorescences

open or dense racemes, axis straight;

buds erect.

dense racemes, axis straight;

buds erect.

Flowers

floral tube 2–10 mm;

sepals reflexed individually or in pairs;

corolla bowl-shaped, petals lavender to purple, purplish red, or dark wine-red, often with red or purple spot near middle, tip, or base, 9–25 mm;

stamens 8, subequal;

ovary 8-grooved, length less than 8 times width;

stigma as long as or exserted beyond anthers.

floral tube 10–15 mm, conspicuously veined, lavender striate within;

sepals reflexed individually;

corolla bowl-shaped, petals lavender shading to white proximally, with large, wedge-shaped purplish red spot near apex, 20–25 mm;

stamens 8, subequal;

ovary 8-grooved, longer than adjacent internode;

stigma exserted beyond anthers.

Capsules

10–30 mm, beak 0–2 mm.

10–15 mm.

Seeds

brown or gray, 1–2 mm, scaly, crest 0.2 mm.

brown or gray, 2 mm, scaly, crest 0.2 mm.

2n

= 16.

Clarkia purpurea

Clarkia imbricata

Phenology Flowering Jun–Jul.
Habitat Clearings, roadsides, chaparral.
Elevation 50 m. (200 ft.)
Distribution
from USDA
w North America; nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Subspecies 3 (3 in the flora).

Clarkia purpurea consists of a diverse assemblage of hexaploid populations and is almost certainly derived from multiple origins followed by hybridization and, perhaps, backcrossing to parental species. Three morphological forms are recognized as subspecies; intergrades are frequent.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Clarkia imbricata, known from only one small area of Sonoma County, is designated as rare by the California Native Plant Society, and is in the Center for Plant Conservation’s National Collection of Endangered Plants; it is a relict of a lineage with 2n = 16 that presumably contributed a genome to the tetraploid species C. davyi and the South American C. tenella.

Morphologically, Clarkia imbricata is most similar to C. speciosa, C. williamsonii, and some populations of C. purpurea. Clarkia imbricata can be distinguished from C. speciosa by the color pattern of the petals and from C. williamsonii and populations of C. purpurea with similar flower size and color pattern by its broader, ascending, overlapping leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades broadly lanceolate to elliptic or ovate; inflorescences dense racemes.
subsp. purpurea
1. Leaf blades linear to lanceolate or narrowly lanceolate; inflorescences usually open racemes.
→ 2
2. Petals 15–25 mm; stigmas exserted beyond anthers.
subsp. viminea
2. Petals 9–14 mm; stigmas not exserted beyond anthers.
subsp. quadrivulnera
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Godetia Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Godetia
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. purpurea subsp. purpurea, C. purpurea subsp. quadrivulnera, C. purpurea subsp. viminea
Synonyms Oenothera purpurea, Godetia purpurea
Name authority (Curtis) A. Nelson & J. F. Macbride: Bot. Gaz. 65: 64. (1918) H. Lewis & M. E. Lewis: Madroño 12: 38. (1953)
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