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purple clarkia, winecup clarkia, winecup fairyfan

arewell-to-spring, farewell-to-spring, farewell-to-spring clarkia, godetia, yellow clarkia

Stems

erect or rarely decumbent, to 100 cm, glabrous and sometimes glaucous or sparsely to densely puberulent, sometimes mixed with longer, spreading hairs.

erect to decumbent, 20–200 cm, puberulent.

Leaves

petiole 0–2 mm;

blade linear or narrowly lanceolate to elliptic or ovate, 1.5–7 cm.

petiole to 10 mm;

blade linear to lanceolate, 1–6 cm.

Inflorescences

open or dense racemes, axis straight;

buds erect.

open or dense spikes or racemes, axis straight;

buds erect.

Flowers

floral tube 2–10 mm;

sepals reflexed individually or in pairs;

corolla bowl-shaped, petals lavender to purple, purplish red, or dark wine-red, often with red or purple spot near middle, tip, or base, 9–25 mm;

stamens 8, subequal;

ovary 8-grooved, length less than 8 times width;

stigma as long as or exserted beyond anthers.

floral tube 3–10 mm;

sepals usually reflexed together to one side, or rarely in pairs or singly;

corolla bowl-shaped, petals pale pink to lavender, usually with red spot or mark near middle, obovate to fan-shaped, 15–60 mm, not lobed, apex sometimes shallowly notched or erose;

stamens 8, in 2 subequal sets;

ovary cylindrical and 4-grooved or fusiform and 8-grooved, puberulent;

stigma exserted or not beyond anthers.

Capsules

10–30 mm, beak 0–2 mm.

15–40 mm, sometimes broader distally;

pedicel 0–13 mm.

Seeds

brown or gray, 1–2 mm, scaly, crest 0.2 mm.

brown to grayish brown, 1–1.5 mm, scaly, crest 0.1 mm.

Clarkia purpurea

Clarkia amoena

Distribution
from USDA
w North America; nw Mexico
[WildflowerSearch map]
[BONAP county map]
from USDA
w North America
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 3 (3 in the flora).

Clarkia purpurea consists of a diverse assemblage of hexaploid populations and is almost certainly derived from multiple origins followed by hybridization and, perhaps, backcrossing to parental species. Three morphological forms are recognized as subspecies; intergrades are frequent.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 5 (5 in the flora).

Clarkia amoena is closely related to C. rubicunda, which differs morphologically mainly in the color pattern of the petals. Petals of C. amoena have a conspicuous red spot or group of small red spots or marks near the middle, whereas those of C. rubicunda have a red area at the base and are not spotted near the middle. The areas of distribution of the two species barely overlap in California just north of San Francisco Bay, and C. rubicunda could be considered a southern geographical race or subspecies of C. amoena were it not that their readily formed hybrids are sterile due to chromosomal rearrangement. Clarkia amoena is one of the parent species of the allotetraploid C. gracilis. Intermediates between subspecies are frequent.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades broadly lanceolate to elliptic or ovate; inflorescences dense racemes.
subsp. purpurea
1. Leaf blades linear to lanceolate or narrowly lanceolate; inflorescences usually open racemes.
→ 2
2. Petals 15–25 mm; stigmas exserted beyond anthers.
subsp. viminea
2. Petals 9–14 mm; stigmas not exserted beyond anthers.
subsp. quadrivulnera
1. Stigmas not exserted beyond anthers; petals less than 20 mm; coastal British Columbia, Oregon, Washington.
subsp. caurina
1. Stigmas exserted beyond anthers; petals 15–60 mm.
→ 2
2. Ovaries 8-grooved, broadly fusiform, 8–12 mm wide.
subsp. whitneyi
2. Ovaries 4-grooved, cylindrical to subclavate, 2–6 mm wide.
→ 3
3. Inflorescences open, internodes longer than subtending flowers, bracts sublinear.
subsp. huntiana
3. Inflorescences congested, internodes shorter than subtending flowers, bracts narrowly lanceolate or wider.
→ 4
4. Stems decumbent to suberect, to 100 cm; petals 20–35 mm, usually with red spot mid blade; coastal California.
subsp. amoena
4. Stems erect, to 200 cm; petals 30–40 mm, usually without red spot or with very small spot or streak mid blade; inland Oregon, Washington.
subsp. lindleyi
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Godetia Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Rhodanthos > subsect. Primigenia
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. purpurea subsp. purpurea, C. purpurea subsp. quadrivulnera, C. purpurea subsp. viminea
C. amoena subsp. amoena, C. amoena subsp. caurina, C. amoena subsp. huntiana, C. amoena subsp. lindleyi, C. amoena subsp. whitneyi
Synonyms Oenothera purpurea, Godetia purpurea Oenothera amoena, Godetia amoena
Name authority (Curtis) A. Nelson & J. F. Macbride: Bot. Gaz. 65: 64. (1918) (Lehmann) A. Nelson & J. F. Macbride: Bot. Gaz. 65: 62. (1918)
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