Flora of North America species comparison

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Clarkia prostrata

Clarkia heterandra

prostrate clarkia

California gaura, heterogaura, mountain clarkia
Stems

prostrate or decumbent, to 50 cm, sparsely puberulent.

erect, to 60 cm, glandular puberulent.
Leaves

sessile or subsessile;

blade oblanceolate to elliptic, 1–2.5 cm, apex usually obtuse.

petiole 5–20 mm;

blade lanceolate to ovate, 2–8 cm.
Inflorescences

prostrate, dense racemes, axis straight;

buds erect.

open racemes or panicles, axis straight;

buds erect.
Flowers

floral tube 4–7 mm;

sepals usually reflexed in pairs;

corolla bowl-shaped, petals lavender-pink shading pale yellow basally, with reddish purple spot above base, 10–15 mm;

stamens 8, subequal;

ovary 8-grooved;

stigma not exserted beyond anthers.

sometimes autogamous;

floral tube 1–2 mm;

sepals reflexed together to 1 side;

corolla rotate, petals elliptic to obovate, tapered to claw;

stamens 8, unequal, anthers cream or light pink, inner ones smaller, sterile;

ovary subglobose, grooves obscure;

stigma not exserted beyond anthers.
Capsules

20–30 mm.

2–3 mm, indehiscent, nutlike;

sessile or pedicellate (to 2 mm).
Seeds

brown or gray, 1–1.5 mm, scaly, crest 0.2 mm.

1 or 2.
2n

= 52.

= 18.

Clarkia prostrata

Clarkia heterandra
Phenology Flowering May–Jul. Flowering May–Jul.
Habitat Coastal bluffs in grasslands and closed-cone pine forests. Shady sites, woodlands, yellow-pine forests.
Elevation 0–100 m. (0–300 ft.) 500–2000 m. (1600–6600 ft.)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Clarkia prostrata, like C. davyi, occurs only on coastal bluffs and adjacent low elevation pine forests along the Pacific coast, and in this case only in the California Central Coast Subregion in Monterey, Santa Barbara, Santa Cruz, and San Luis Obispo counties.

Clarkia prostrata is a hexaploid that combines the tetraploid genome of C. davyi and the diploid genome of C. speciosa. Clarkia prostrata is morphologically and ecologically very similar to C. davyi but can usually be distinguished by its larger flowers with a spot on each petal. It differs from C. speciosa by having smaller flowers with the stigma not exserted beyond the anthers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Clarkia heterandra occurs in southern Oregon and California, where it is known from the Klamath Ranges in Trinity County, widely in the Sierra Nevada and Tehachapi Mountain area from Nevada to Kern counties, and in the South Coast and Transverse Ranges from San Luis Obispo to Riverside and San Bernardino counties.

Prior to 1986, Clarkia heterandra was treated as the monotypic genus Heterogaura, thought to be possibly related to the genus Gaura (now a section of Oenothera) due to its similar indehiscent fruits. However, molecular data indicate that C. heterandra is clearly within Clarkia, most closely related to C. dudleyana and C. unguiculata, which suggests a possible hybrid origin at the diploid level (K. J. Sytsma and L. D. Gottlieb 1986, 1986b; W. L. Wagner et al. 2007). Clarkia dudleyana and C. unguiculata produce spontaneous hybrids when grown adjacent to each other if they come from allopatric populations.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Godetia Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Phaeostoma > subsect. Heterogaura
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Synonyms Gaura heterandra, Heterogaura heterandra
Name authority H. Lewis & M. E. Lewis: Madroño 12: 36. (1953) (Torrey) H. Lewis & P. H. Raven: Madroño 39: 163. (1992)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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