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prostrate clarkia

arewell-to-spring, farewell-to-spring, farewell-to-spring clarkia, godetia, yellow clarkia

Stems

prostrate or decumbent, to 50 cm, sparsely puberulent.

erect to decumbent, 20–200 cm, puberulent.

Leaves

sessile or subsessile;

blade oblanceolate to elliptic, 1–2.5 cm, apex usually obtuse.

petiole to 10 mm;

blade linear to lanceolate, 1–6 cm.

Inflorescences

prostrate, dense racemes, axis straight;

buds erect.

open or dense spikes or racemes, axis straight;

buds erect.

Flowers

floral tube 4–7 mm;

sepals usually reflexed in pairs;

corolla bowl-shaped, petals lavender-pink shading pale yellow basally, with reddish purple spot above base, 10–15 mm;

stamens 8, subequal;

ovary 8-grooved;

stigma not exserted beyond anthers.

floral tube 3–10 mm;

sepals usually reflexed together to one side, or rarely in pairs or singly;

corolla bowl-shaped, petals pale pink to lavender, usually with red spot or mark near middle, obovate to fan-shaped, 15–60 mm, not lobed, apex sometimes shallowly notched or erose;

stamens 8, in 2 subequal sets;

ovary cylindrical and 4-grooved or fusiform and 8-grooved, puberulent;

stigma exserted or not beyond anthers.

Capsules

20–30 mm.

15–40 mm, sometimes broader distally;

pedicel 0–13 mm.

Seeds

brown or gray, 1–1.5 mm, scaly, crest 0.2 mm.

brown to grayish brown, 1–1.5 mm, scaly, crest 0.1 mm.

2n

= 52.

Clarkia prostrata

Clarkia amoena

Phenology Flowering May–Jul.
Habitat Coastal bluffs in grasslands and closed-cone pine forests.
Elevation 0–100 m. (0–300 ft.)
Distribution
from FNA
CA
[BONAP county map]
from USDA
w North America
[WildflowerSearch map]
[BONAP county map]
Discussion

Clarkia prostrata, like C. davyi, occurs only on coastal bluffs and adjacent low elevation pine forests along the Pacific coast, and in this case only in the California Central Coast Subregion in Monterey, Santa Barbara, Santa Cruz, and San Luis Obispo counties.

Clarkia prostrata is a hexaploid that combines the tetraploid genome of C. davyi and the diploid genome of C. speciosa. Clarkia prostrata is morphologically and ecologically very similar to C. davyi but can usually be distinguished by its larger flowers with a spot on each petal. It differs from C. speciosa by having smaller flowers with the stigma not exserted beyond the anthers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 5 (5 in the flora).

Clarkia amoena is closely related to C. rubicunda, which differs morphologically mainly in the color pattern of the petals. Petals of C. amoena have a conspicuous red spot or group of small red spots or marks near the middle, whereas those of C. rubicunda have a red area at the base and are not spotted near the middle. The areas of distribution of the two species barely overlap in California just north of San Francisco Bay, and C. rubicunda could be considered a southern geographical race or subspecies of C. amoena were it not that their readily formed hybrids are sterile due to chromosomal rearrangement. Clarkia amoena is one of the parent species of the allotetraploid C. gracilis. Intermediates between subspecies are frequent.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stigmas not exserted beyond anthers; petals less than 20 mm; coastal British Columbia, Oregon, Washington.
subsp. caurina
1. Stigmas exserted beyond anthers; petals 15–60 mm.
→ 2
2. Ovaries 8-grooved, broadly fusiform, 8–12 mm wide.
subsp. whitneyi
2. Ovaries 4-grooved, cylindrical to subclavate, 2–6 mm wide.
→ 3
3. Inflorescences open, internodes longer than subtending flowers, bracts sublinear.
subsp. huntiana
3. Inflorescences congested, internodes shorter than subtending flowers, bracts narrowly lanceolate or wider.
→ 4
4. Stems decumbent to suberect, to 100 cm; petals 20–35 mm, usually with red spot mid blade; coastal California.
subsp. amoena
4. Stems erect, to 200 cm; petals 30–40 mm, usually without red spot or with very small spot or streak mid blade; inland Oregon, Washington.
subsp. lindleyi
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Godetia Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Rhodanthos > subsect. Primigenia
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. amoena subsp. amoena, C. amoena subsp. caurina, C. amoena subsp. huntiana, C. amoena subsp. lindleyi, C. amoena subsp. whitneyi
Synonyms Oenothera amoena, Godetia amoena
Name authority H. Lewis & M. E. Lewis: Madroño 12: 36. (1953) (Lehmann) A. Nelson & J. F. Macbride: Bot. Gaz. 65: 62. (1918)
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