Clarkia mildrediae |
Clarkia xantiana |
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Mildred's clarkia |
gunsight clarkia, Xantus' clarkia |
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Stems | erect, to 100 cm, puberulent. |
erect, to 80 cm, glabrous, glaucous. |
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Leaves | petiole 15–40 mm; blade elliptic to ovate, 3–6 cm. |
petiole 0–2 mm; blade linear to lanceolate, 2–6 cm. |
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Inflorescences | open racemes, axis recurved in bud, straight only 1–3 nodes distal to open flowers; buds pendent, tip acute. |
open racemes, axis straight; buds pendent. |
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Flowers | floral tube 2–3 mm; sepals usually reflexed individually; corolla rotate, petals reddish purple, often darker flecked or spotted, narrowly obdeltate to suborbiculate, unlobed, 11–25 × 7–18 mm, length 1.4–1.6 times width; stamens 8, subequal, subtended by ciliate scales, anthers magenta or orange-red to yellow, pollen blue-gray or bright yellow to tan; ovary shallowly 4-grooved, puberulent; stigma exserted beyond anthers. |
floral tube 2–5 mm; sepals reflexed together to 1 side; corolla rotate, petals lavender to reddish purple, lavender-pink, or white, often with white-surrounded dark reddish purple spot distally, clawed, 2-lobed, with slender central tooth, 1–3 mm, 6–20 mm; stamens 8, in 2 unequal sets, outer anthers lavender to purple, inner smaller, paler; ovary 8-grooved; stigma exserted or not beyond anthers. |
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Capsules | 20–30 mm; pedicel 0–5 mm. |
15–25 mm; pedicel 0–5 mm. |
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Seeds | brown or gray, 1.5–1.8 mm, scaly-echinate, crest 0.1 mm. |
brown, 1.3–1.5 mm, tuberculate, crest minute. |
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Clarkia mildrediae |
Clarkia xantiana |
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Distribution |
California
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California
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Discussion | Subspecies 2 (2 in the flora). Clarkia mildrediae is closely related to, and may be derived from, C. borealis. Clarkia mildrediae subsp. lutescens is probably the direct ancestor of C. stellata. Clarkia mildrediae differs from C. borealis in petal color and inflorescence habit and from C. stellata in flower size and position of the stigma. On the basis of morphology, chromosome number and pairing, C. mildrediae appears to be one of the parents of the tetraploid species C. rhomboidea. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 2 (2 in the flora). Originally placed in the group now delimited as subsect. Phaeostoma by H. Lewis and M. E. Lewis (1955), Clarkia xantiana was treated as a monotypic subsect. Xantianae within sect. Phaeostoma by K. E. Holsinger (1985), based mainly on its unusual 2-lobed petals with a tooth in the sinus. Molecular data (R. A. Levin et al. 2004) placed C. xantiana close to C. bottae, and both of them close to but not within sect. Phaeostoma. Both species share the chromosome number 2n = 18 with C. jolonensis. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 10. | FNA vol. 10. | ||||||||
Parent taxa | Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Myxocarpa | Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Fibula | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Phaeostoma mildrediae | |||||||||
Name authority | (A. Heller) H. Lewis & M. E. Lewis: Madroño 12: 34. (1953) | A. Gray: Proc. Boston Soc. Nat. Hist. 7: 146. (1859) | ||||||||
Web links |