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Mildred's clarkia

red spotted clarkia, red spotted or redspot clarkia, redspot clarkia

Stems

erect, to 100 cm, puberulent.

erect or decumbent, to 60 cm, puberulent.

Leaves

petiole 15–40 mm;

blade elliptic to ovate, 3–6 cm.

petiole 0–5 mm;

blade linear to lanceolate, 1–6 cm.

Inflorescences

open racemes, axis recurved in bud, straight only 1–3 nodes distal to open flowers;

buds pendent, tip acute.

open or dense racemes or panicles, axis straight;

buds erect.

Flowers

floral tube 2–3 mm;

sepals usually reflexed individually;

corolla rotate, petals reddish purple, often darker flecked or spotted, narrowly obdeltate to suborbiculate, unlobed, 11–25 × 7–18 mm, length 1.4–1.6 times width;

stamens 8, subequal, subtended by ciliate scales, anthers magenta or orange-red to yellow, pollen blue-gray or bright yellow to tan;

ovary shallowly 4-grooved, puberulent;

stigma exserted beyond anthers.

floral tube 5–15 mm;

sepals usually reflexed individually or in pairs, or rarely together to one side, tips not distinct in bud;

corolla bowl-shaped, petals lavender to wine-red, often shading white or pale yellow toward base, unspotted or with bright red or purplish red spot near or proximal to middle, 10–25 mm;

stamens 8, subequal;

ovary 8-grooved;

stigma exserted beyond anthers.

Capsules

20–30 mm;

pedicel 0–5 mm.

15–20 mm.

Seeds

brown or gray, 1.5–1.8 mm, scaly-echinate, crest 0.1 mm.

brown or gray, 0.7–1 mm, scaly, crest 0.1 mm.

Clarkia mildrediae

Clarkia speciosa

Distribution
from USDA
California
[WildflowerSearch map]
[BONAP county map]
from USDA
California
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Clarkia mildrediae is closely related to, and may be derived from, C. borealis. Clarkia mildrediae subsp. lutescens is probably the direct ancestor of C. stellata. Clarkia mildrediae differs from C. borealis in petal color and inflorescence habit and from C. stellata in flower size and position of the stigma. On the basis of morphology, chromosome number and pairing, C. mildrediae appears to be one of the parents of the tetraploid species C. rhomboidea.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 4 (4 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals 12–18 mm wide; anthers magenta, fresh pollen blue-gray.
subsp. mildrediae
1. Petals 7–16 mm wide; anthers yellow to orange-red, fresh pollen bright yellow to tan.
subsp. lutescens
1. Petals without red or purplish red spot.
→ 2
2. Stems erect to decumbent; petals dark purplish red to lavender, ± white or pale yellow near base, rarely pale yellow throughout.
subsp. speciosa
2. Stems decumbent; petals lavender to pink shading pale yellow or white in proximal 1/2.
subsp. immaculata
1. Petals with bright red or purplish red spot near or proximal to middle.
→ 3
3. Inflorescences dense racemes.
subsp. nitens
3. Inflorescences open racemes or panicles.
→ 4
4. Petals usually with bright red spot; branches on well-developed plants many, few-flowered.
subsp. speciosa
4. Petals with purplish red spot; branches on well-developed plants virgate, few, many- flowered.
subsp. polyantha
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Myxocarpa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Godetia
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. mildrediae subsp. lutescens, C. mildrediae subsp. mildrediae
C. speciosa subsp. immaculata, C. speciosa subsp. nitens, C. speciosa subsp. polyantha, C. speciosa subsp. speciosa
Synonyms Phaeostoma mildrediae Oenothera viminea var. parviflora, Godetia parviflora
Name authority (A. Heller) H. Lewis & M. E. Lewis: Madroño 12: 34. (1953) H. Lewis & M. E. Lewis: Madroño 12: 34. (1953)
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