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Mildred's clarkia

Ramona clarkia

Stems

erect, to 100 cm, puberulent.

erect, 30–90 cm, puberulent.

Leaves

petiole 15–40 mm;

blade elliptic to ovate, 3–6 cm.

petiole to 8 mm;

blade narrowly lanceolate to elliptic, 2–4 cm.

Inflorescences

open racemes, axis recurved in bud, straight only 1–3 nodes distal to open flowers;

buds pendent, tip acute.

open racemes, axis recurved at tip in bud;

buds pendent.

Flowers

floral tube 2–3 mm;

sepals usually reflexed individually;

corolla rotate, petals reddish purple, often darker flecked or spotted, narrowly obdeltate to suborbiculate, unlobed, 11–25 × 7–18 mm, length 1.4–1.6 times width;

stamens 8, subequal, subtended by ciliate scales, anthers magenta or orange-red to yellow, pollen blue-gray or bright yellow to tan;

ovary shallowly 4-grooved, puberulent;

stigma exserted beyond anthers.

floral tube 1.5–2 mm;

sepals reflexed together to 1 side;

corolla rotate to bowl-shaped, petals pale pink shading nearly white near base, purple-flecked, fading pink, oblanceolate or obovate to diamond-shaped, 6–10 mm;

stamens 8, unequal, outer anthers dark pink, inner smaller, paler;

ovary shallowly 8-grooved.

Capsules

20–30 mm;

pedicel 0–5 mm.

15–30 mm;

subsessile.

Seeds

brown or gray, 1.5–1.8 mm, scaly-echinate, crest 0.1 mm.

brown, 1 mm, tuberculate, crest inconspicuous.

2n

= 34.

Clarkia mildrediae

Clarkia similis

Phenology Flowering Apr–Jun.
Habitat Shady sites, oak woodlands, chaparral.
Elevation 0–1000 m. (0–3300 ft.)
Distribution
from USDA
California
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Clarkia mildrediae is closely related to, and may be derived from, C. borealis. Clarkia mildrediae subsp. lutescens is probably the direct ancestor of C. stellata. Clarkia mildrediae differs from C. borealis in petal color and inflorescence habit and from C. stellata in flower size and position of the stigma. On the basis of morphology, chromosome number and pairing, C. mildrediae appears to be one of the parents of the tetraploid species C. rhomboidea.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Clarkia similis is known from southwestern California and northern Baja California, Mexico, mainly in the South Coast, Transverse, and Peninsular ranges, from Monterey and San Benito counties in the north to San Diego County in the south. Reports from the Southern Sierra Nevada Foothills in Kern County and the Sacramento Valley in Tehama County are not confirmed.

Clarkia similis is a tetraploid species derived through polyploidization following hybridization between C. epilobioides and C. modesta.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals 12–18 mm wide; anthers magenta, fresh pollen blue-gray.
subsp. mildrediae
1. Petals 7–16 mm wide; anthers yellow to orange-red, fresh pollen bright yellow to tan.
subsp. lutescens
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Myxocarpa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Phaeostoma > subsect. Prognatae
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. mildrediae subsp. lutescens, C. mildrediae subsp. mildrediae
Synonyms Phaeostoma mildrediae
Name authority (A. Heller) H. Lewis & M. E. Lewis: Madroño 12: 34. (1953) H. Lewis & W. R. Ernst: Madroño 12: 89. (1953)
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