Flora of North America species comparison

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Clarkia mildrediae

Clarkia rubicunda

Mildred's clarkia

farewell-to-spring, ruby chalice clarkia
Stems

erect, to 100 cm, puberulent.

erect or decumbent, to 150 cm, puberulent;

buds erect.
Leaves

petiole 15–40 mm;

blade elliptic to ovate, 3–6 cm.

petiole to 10 mm;

blade lanceolate to elliptic, 1–4 cm.
Inflorescences

open racemes, axis recurved in bud, straight only 1–3 nodes distal to open flowers;

buds pendent, tip acute.

open or dense spikes or racemes, axis straight;

buds erect.
Flowers

floral tube 2–3 mm;

sepals usually reflexed individually;

corolla rotate, petals reddish purple, often darker flecked or spotted, narrowly obdeltate to suborbiculate, unlobed, 11–25 × 7–18 mm, length 1.4–1.6 times width;

stamens 8, subequal, subtended by ciliate scales, anthers magenta or orange-red to yellow, pollen blue-gray or bright yellow to tan;

ovary shallowly 4-grooved, puberulent;

stigma exserted beyond anthers.

floral tube 4–10 mm;

sepals reflexed together to one side;

corolla bowl-shaped, petals pink to lavender, base red or purplish red, fan-shaped, 10–30 mm, apex erose;

stamens 8, subequal;

ovary cylindrical, 4-grooved, puberulent;

stigma exserted beyond anthers.
Capsules

20–30 mm;

pedicel 0–5 mm.

20–40 mm;

pedicel 0–25(–40) mm.
Seeds

brown or gray, 1.5–1.8 mm, scaly-echinate, crest 0.1 mm.

brown to grayish brown, 1.2–1.5 mm, scaly, crest 0.2 mm.
2n

= 14.

Clarkia mildrediae

Clarkia rubicunda
Phenology Flowering May–Aug.
Habitat Openings in woodlands, forests, chaparral, coastal scrub.
Elevation 0–500 m. (0–1600 ft.)
Distribution
from USDA
California
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Clarkia mildrediae is closely related to, and may be derived from, C. borealis. Clarkia mildrediae subsp. lutescens is probably the direct ancestor of C. stellata. Clarkia mildrediae differs from C. borealis in petal color and inflorescence habit and from C. stellata in flower size and position of the stigma. On the basis of morphology, chromosome number and pairing, C. mildrediae appears to be one of the parents of the tetraploid species C. rhomboidea.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Clarkia rubicunda is known from the central coast of California, from Contra Costa and Marin counties south along the coast and foothills to northern San Luis Obispo County.

Clarkia rubicunda is probably a derivative of C. amoena and may be ancestral to C. franciscana. Clarkia rubicunda is distinguishable from some populations of C. amoena only by the absence of a red spot or group of spots near the middle of the petal and the presence of a red area at the base of the petal. Clarkia rubicunda can be distinguished from C. franciscana by the position of the stigma and size and shape of the petals. All three species differ in chromosome arrangement and hybrids are highly sterile.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key
1. Petals 12–18 mm wide; anthers magenta, fresh pollen blue-gray.
subsp. mildrediae
1. Petals 7–16 mm wide; anthers yellow to orange-red, fresh pollen bright yellow to tan.
subsp. lutescens
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Myxocarpa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Rhodanthos > subsect. Primigenia
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. mildrediae subsp. lutescens, C. mildrediae subsp. mildrediae
Synonyms Phaeostoma mildrediae Godetia rubicunda, C. rubicunda subsp. blasdalei, G. blasdalei
Name authority (A. Heller) H. Lewis & M. E. Lewis: Madroño 12: 34. (1953) (Lindley) H. Lewis & M. E. Lewis: Madroño 12: 34. (1953)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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