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Mildred's clarkia

beak clarkia

Stems

erect, to 100 cm, puberulent.

erect, to 60 cm, puberulent.

Leaves

petiole 15–40 mm;

blade elliptic to ovate, 3–6 cm.

petiole to 10 mm;

blade lanceolate, 1–6 cm.

Inflorescences

open racemes, axis recurved in bud, straight only 1–3 nodes distal to open flowers;

buds pendent, tip acute.

open racemes, axis recurved at tip in bud;

buds pendent.

Flowers

floral tube 2–3 mm;

sepals usually reflexed individually;

corolla rotate, petals reddish purple, often darker flecked or spotted, narrowly obdeltate to suborbiculate, unlobed, 11–25 × 7–18 mm, length 1.4–1.6 times width;

stamens 8, subequal, subtended by ciliate scales, anthers magenta or orange-red to yellow, pollen blue-gray or bright yellow to tan;

ovary shallowly 4-grooved, puberulent;

stigma exserted beyond anthers.

floral tube 1.5–2.5 mm, with ring of hairs at distal margin inside;

sepals reflexed together to 1 side;

corolla bowl-shaped, petals pinkish lavender shading white near middle, often flecked reddish purple, base reddish purple, 10–25 mm;

stamens 8, unequal, width of all filaments equal or inner slightly thinner, outer anthers lavender, inner smaller, paler.

Capsules

20–30 mm;

pedicel 0–5 mm.

10–30 mm, beak 7–15 mm.

Seeds

brown or gray, 1.5–1.8 mm, scaly-echinate, crest 0.1 mm.

unknown.

2n

= 18.

Clarkia mildrediae

Clarkia rostrata

Phenology Flowering Apr–May.
Habitat Oak-pine woodlands.
Elevation 500 m. (1600 ft.)
Distribution
from USDA
California
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Clarkia mildrediae is closely related to, and may be derived from, C. borealis. Clarkia mildrediae subsp. lutescens is probably the direct ancestor of C. stellata. Clarkia mildrediae differs from C. borealis in petal color and inflorescence habit and from C. stellata in flower size and position of the stigma. On the basis of morphology, chromosome number and pairing, C. mildrediae appears to be one of the parents of the tetraploid species C. rhomboidea.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Clarkia rostrata is known only from the Merced River drainage in the central Sierra Nevada Foothills, including Mariposa, Merced, Stanislaus, and (barely) Tuolumne counties. Because of its very limited distribution, C. rostrata is listed as rare by the California Native Plant Society.

Clarkia rostrata is closely related to C. cylindrica and C. lewisii but can be distinguished readily from both by the conspicuous beak of the capsule.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals 12–18 mm wide; anthers magenta, fresh pollen blue-gray.
subsp. mildrediae
1. Petals 7–16 mm wide; anthers yellow to orange-red, fresh pollen bright yellow to tan.
subsp. lutescens
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Myxocarpa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Phaeostoma > subsect. Sympherica
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. mildrediae subsp. lutescens, C. mildrediae subsp. mildrediae
Synonyms Phaeostoma mildrediae
Name authority (A. Heller) H. Lewis & M. E. Lewis: Madroño 12: 34. (1953) W. S. Davis: Brittonia 22: 281. (1970)
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