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Mildred's clarkia

California gaura, heterogaura, mountain clarkia

Stems

erect, to 100 cm, puberulent.

erect, to 60 cm, glandular puberulent.

Leaves

petiole 15–40 mm;

blade elliptic to ovate, 3–6 cm.

petiole 5–20 mm;

blade lanceolate to ovate, 2–8 cm.

Inflorescences

open racemes, axis recurved in bud, straight only 1–3 nodes distal to open flowers;

buds pendent, tip acute.

open racemes or panicles, axis straight;

buds erect.

Flowers

floral tube 2–3 mm;

sepals usually reflexed individually;

corolla rotate, petals reddish purple, often darker flecked or spotted, narrowly obdeltate to suborbiculate, unlobed, 11–25 × 7–18 mm, length 1.4–1.6 times width;

stamens 8, subequal, subtended by ciliate scales, anthers magenta or orange-red to yellow, pollen blue-gray or bright yellow to tan;

ovary shallowly 4-grooved, puberulent;

stigma exserted beyond anthers.

sometimes autogamous;

floral tube 1–2 mm;

sepals reflexed together to 1 side;

corolla rotate, petals elliptic to obovate, tapered to claw;

stamens 8, unequal, anthers cream or light pink, inner ones smaller, sterile;

ovary subglobose, grooves obscure;

stigma not exserted beyond anthers.

Capsules

20–30 mm;

pedicel 0–5 mm.

2–3 mm, indehiscent, nutlike;

sessile or pedicellate (to 2 mm).

Seeds

brown or gray, 1.5–1.8 mm, scaly-echinate, crest 0.1 mm.

1 or 2.

2n

= 18.

Clarkia mildrediae

Clarkia heterandra

Phenology Flowering May–Jul.
Habitat Shady sites, woodlands, yellow-pine forests.
Elevation 500–2000 m. (1600–6600 ft.)
Distribution
from USDA
California
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Clarkia mildrediae is closely related to, and may be derived from, C. borealis. Clarkia mildrediae subsp. lutescens is probably the direct ancestor of C. stellata. Clarkia mildrediae differs from C. borealis in petal color and inflorescence habit and from C. stellata in flower size and position of the stigma. On the basis of morphology, chromosome number and pairing, C. mildrediae appears to be one of the parents of the tetraploid species C. rhomboidea.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Clarkia heterandra occurs in southern Oregon and California, where it is known from the Klamath Ranges in Trinity County, widely in the Sierra Nevada and Tehachapi Mountain area from Nevada to Kern counties, and in the South Coast and Transverse Ranges from San Luis Obispo to Riverside and San Bernardino counties.

Prior to 1986, Clarkia heterandra was treated as the monotypic genus Heterogaura, thought to be possibly related to the genus Gaura (now a section of Oenothera) due to its similar indehiscent fruits. However, molecular data indicate that C. heterandra is clearly within Clarkia, most closely related to C. dudleyana and C. unguiculata, which suggests a possible hybrid origin at the diploid level (K. J. Sytsma and L. D. Gottlieb 1986, 1986b; W. L. Wagner et al. 2007). Clarkia dudleyana and C. unguiculata produce spontaneous hybrids when grown adjacent to each other if they come from allopatric populations.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals 12–18 mm wide; anthers magenta, fresh pollen blue-gray.
subsp. mildrediae
1. Petals 7–16 mm wide; anthers yellow to orange-red, fresh pollen bright yellow to tan.
subsp. lutescens
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Myxocarpa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Phaeostoma > subsect. Heterogaura
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. mildrediae subsp. lutescens, C. mildrediae subsp. mildrediae
Synonyms Phaeostoma mildrediae Gaura heterandra, Heterogaura heterandra
Name authority (A. Heller) H. Lewis & M. E. Lewis: Madroño 12: 34. (1953) (Torrey) H. Lewis & P. H. Raven: Madroño 39: 163. (1992)
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