Flora of North America species comparison

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Clarkia mildrediae

Clarkia dudleyana

Mildred's clarkia

Dudley's clarkia
Stems

erect, to 100 cm, puberulent.

erect, to 70 cm, puberulent.
Leaves

petiole 15–40 mm;

blade elliptic to ovate, 3–6 cm.

petiole 3–10 mm;

blade narrowly lanceolate, 1.5–7 cm.
Inflorescences

open racemes, axis recurved in bud, straight only 1–3 nodes distal to open flowers;

buds pendent, tip acute.

open racemes, axis recurved at tip in bud;

buds pendent.
Flowers

floral tube 2–3 mm;

sepals usually reflexed individually;

corolla rotate, petals reddish purple, often darker flecked or spotted, narrowly obdeltate to suborbiculate, unlobed, 11–25 × 7–18 mm, length 1.4–1.6 times width;

stamens 8, subequal, subtended by ciliate scales, anthers magenta or orange-red to yellow, pollen blue-gray or bright yellow to tan;

ovary shallowly 4-grooved, puberulent;

stigma exserted beyond anthers.

floral tube 1–3 mm;

sepals reflexed together to 1 side;

corolla bowl-shaped, petals lavender-pink, usually white-streaked, often red-flecked, broadly fan-shaped, 10–30 mm, apex subentire to crenulate;

stamens 8, unequal, outer anthers lavender, inner smaller, paler.
Capsules

20–30 mm;

pedicel 0–5 mm.

10–30 mm.
Seeds

brown or gray, 1.5–1.8 mm, scaly-echinate, crest 0.1 mm.

brown, 1 mm, minutely scaly to puberulent, crest inconspicuous.
2n

= 18.

Clarkia mildrediae

Clarkia dudleyana
Phenology Flowering May–Jul.
Habitat Openings in woodlands, chaparral, yellow-pine forests, coastal sage.
Elevation 0–1500 m. (0–4900 ft.)
Distribution
from USDA
California
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Clarkia mildrediae is closely related to, and may be derived from, C. borealis. Clarkia mildrediae subsp. lutescens is probably the direct ancestor of C. stellata. Clarkia mildrediae differs from C. borealis in petal color and inflorescence habit and from C. stellata in flower size and position of the stigma. On the basis of morphology, chromosome number and pairing, C. mildrediae appears to be one of the parents of the tetraploid species C. rhomboidea.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Clarkia dudleyana is a rather widespread species in California, known primarily from the central and southern Sierra Nevada foothills, the Tehachapi Mountain area, the Transverse Ranges, and the Peninsular Ranges, ranging from Tuolumne to Riverside counties, sporadically in the north to Nevada County and in the south to San Diego County

Clarkia dudleyana is morphologically most similar to C. biloba and C. modesta, but molecular data suggest that the relationship is not close. On the basis of chloroplast DNA sequence, C. dudleyana and C. heterandra are closely related.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key
1. Petals 12–18 mm wide; anthers magenta, fresh pollen blue-gray.
subsp. mildrediae
1. Petals 7–16 mm wide; anthers yellow to orange-red, fresh pollen bright yellow to tan.
subsp. lutescens
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Myxocarpa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Phaeostoma > subsect. Lautiflorae
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. mildrediae subsp. lutescens, C. mildrediae subsp. mildrediae
Synonyms Phaeostoma mildrediae Godetia dudleyana
Name authority (A. Heller) H. Lewis & M. E. Lewis: Madroño 12: 34. (1953) (Abrams) J. F. Macbride: Contr. Gray Herb. 56: 54. (1918)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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