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Mildred's clarkia

Brewer's clarkia, fairy fans

Stems

erect, to 100 cm, puberulent.

erect or decumbent, to 20 cm, glabrous or sparsely puberulent.

Leaves

petiole 15–40 mm;

blade elliptic to ovate, 3–6 cm.

petiole to 20 mm;

blade linear to lanceolate, 2–5 cm.

Inflorescences

open racemes, axis recurved in bud, straight only 1–3 nodes distal to open flowers;

buds pendent, tip acute.

racemes, axis straight or recurved;

buds pendent.

Flowers

floral tube 2–3 mm;

sepals usually reflexed individually;

corolla rotate, petals reddish purple, often darker flecked or spotted, narrowly obdeltate to suborbiculate, unlobed, 11–25 × 7–18 mm, length 1.4–1.6 times width;

stamens 8, subequal, subtended by ciliate scales, anthers magenta or orange-red to yellow, pollen blue-gray or bright yellow to tan;

ovary shallowly 4-grooved, puberulent;

stigma exserted beyond anthers.

floral tube 20–35 mm;

sepals reflexed together to 1 side, not petal-like, green to magenta;

corolla rotate, petals pink, broadly fan-shaped, 15–25 mm, length equal to width, without claw, 3-lobed, middle lobe longer and much narrower, linear to oblanceolate;

stamens 4, filaments wider distally;

ovary inconspicuously grooved;

stigma exserted beyond anthers.

Capsules

20–30 mm;

pedicel 0–5 mm.

15–40 mm;

subsessile.

Seeds

brown or gray, 1.5–1.8 mm, scaly-echinate, crest 0.1 mm.

reddish brown, 2–3 mm, scaly-tuberculate, crest to 0.8 mm, conspicuous.

2n

= 14.

Clarkia mildrediae

Clarkia breweri

Phenology Flowering Apr–Jun.
Habitat Woodlands, chaparral.
Elevation 0–1000 m. (0–3300 ft.)
Distribution
from USDA
California
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Clarkia mildrediae is closely related to, and may be derived from, C. borealis. Clarkia mildrediae subsp. lutescens is probably the direct ancestor of C. stellata. Clarkia mildrediae differs from C. borealis in petal color and inflorescence habit and from C. stellata in flower size and position of the stigma. On the basis of morphology, chromosome number and pairing, C. mildrediae appears to be one of the parents of the tetraploid species C. rhomboidea.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Clarkia breweri is restricted to dry woodlands and chaparral west of the Central Valley from the San Francisco Bay area into the southern Coast Ranges in Fresno, Monterey, and San Benito counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals 12–18 mm wide; anthers magenta, fresh pollen blue-gray.
subsp. mildrediae
1. Petals 7–16 mm wide; anthers yellow to orange-red, fresh pollen bright yellow to tan.
subsp. lutescens
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Myxocarpa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Eucharidium
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. mildrediae subsp. lutescens, C. mildrediae subsp. mildrediae
Synonyms Phaeostoma mildrediae Eucharidium breweri
Name authority (A. Heller) H. Lewis & M. E. Lewis: Madroño 12: 34. (1953) (A. Gray) Greene: Pittonia 1: 141. (1887)
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