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Mildred's clarkia

botta's clarkia, punch bowl godetia

Stems

erect, to 100 cm, puberulent.

erect, to 100 cm, glabrous, glaucous, rarely puberulent.

Leaves

petiole 15–40 mm;

blade elliptic to ovate, 3–6 cm.

petiole to 5 mm;

blade narrowly lanceolate to lanceolate, 3–10 cm.

Inflorescences

open racemes, axis recurved in bud, straight only 1–3 nodes distal to open flowers;

buds pendent, tip acute.

open racemes, axis straight in bud;

buds pendent.

Flowers

floral tube 2–3 mm;

sepals usually reflexed individually;

corolla rotate, petals reddish purple, often darker flecked or spotted, narrowly obdeltate to suborbiculate, unlobed, 11–25 × 7–18 mm, length 1.4–1.6 times width;

stamens 8, subequal, subtended by ciliate scales, anthers magenta or orange-red to yellow, pollen blue-gray or bright yellow to tan;

ovary shallowly 4-grooved, puberulent;

stigma exserted beyond anthers.

floral tube 2–3 mm;

sepals reflexed together to 1 side;

corolla bowl-shaped, petals pale lavender to pinkish lavender, often white toward base, usually red-flecked, unlobed, 15–30 mm;

stamens 8, unequal, outer anthers lavender, inner smaller, paler;

ovary obscurely 4-grooved;

stigma exserted beyond anthers.

Capsules

20–30 mm;

pedicel 0–5 mm.

30–40 mm;

pedicel 0–30 mm.

Seeds

brown or gray, 1.5–1.8 mm, scaly-echinate, crest 0.1 mm.

brown or gray, 1.2–1.8 mm, ± papillose, crest 0.2 mm.

2n

= 18.

Clarkia mildrediae

Clarkia bottae

Phenology Flowering Apr–Jul.
Habitat Chaparral, woodlands, coastal scrub.
Elevation 0–1000 m. (0–3300 ft.)
Distribution
from USDA
California
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Clarkia mildrediae is closely related to, and may be derived from, C. borealis. Clarkia mildrediae subsp. lutescens is probably the direct ancestor of C. stellata. Clarkia mildrediae differs from C. borealis in petal color and inflorescence habit and from C. stellata in flower size and position of the stigma. On the basis of morphology, chromosome number and pairing, C. mildrediae appears to be one of the parents of the tetraploid species C. rhomboidea.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Clarkia bottae is known from the Outer South Coast Ranges from Monterey and San Benito to Santa Barbara counties, throughout southwestern California, and more sporadically in the southern Sierra Nevada in Fresno and Kern counties. Reports from Del Norte, Madera, and Napa counties have not been verified.

Clarkia bottae is most closely related to C. jolonensis and may be closely related to C. xantiana; all three have 2n = 18, and spontaneous hybrids were formed when C. bottae was grown adjacent to a natural population of C. xantiana.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals 12–18 mm wide; anthers magenta, fresh pollen blue-gray.
subsp. mildrediae
1. Petals 7–16 mm wide; anthers yellow to orange-red, fresh pollen bright yellow to tan.
subsp. lutescens
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Myxocarpa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Fibula
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. mildrediae subsp. lutescens, C. mildrediae subsp. mildrediae
Synonyms Phaeostoma mildrediae Godetia bottae, C. deflexa, G. bottae var. deflexa, G. deflexa
Name authority (A. Heller) H. Lewis & M. E. Lewis: Madroño 12: 34. (1953) (Spach) H. Lewis & M. E. Lewis: Madroño 12: 33. (1953)
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