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Mildred's clarkia

glandular clarkia, glandular clarkia or fairyfan, glandular fairyfan

Stems

erect, to 100 cm, puberulent.

erect, to 8 cm, glabrous or puberulent.

Leaves

petiole 15–40 mm;

blade elliptic to ovate, 3–6 cm.

sessile;

blade linear to narrowly lanceolate or oblanceolate, 1.5–6cm, surfaces glabrate.

Inflorescences

open racemes, axis recurved in bud, straight only 1–3 nodes distal to open flowers;

buds pendent, tip acute.

racemes, axis recurved at tip in bud;

buds pendent.

Flowers

floral tube 2–3 mm;

sepals usually reflexed individually;

corolla rotate, petals reddish purple, often darker flecked or spotted, narrowly obdeltate to suborbiculate, unlobed, 11–25 × 7–18 mm, length 1.4–1.6 times width;

stamens 8, subequal, subtended by ciliate scales, anthers magenta or orange-red to yellow, pollen blue-gray or bright yellow to tan;

ovary shallowly 4-grooved, puberulent;

stigma exserted beyond anthers.

floral tube 3–7 mm, puberulent with spreading hairs and shorter glandular hairs;

corolla bowl-shaped, petals pinkish lavender, lighter proximally, often with dark reddish spot at base, broadly obovate or obdeltate, 10–30 mm;

stamens 8, subequal;

ovary 8-grooved, sparsely puberulent, hairs mostly glandular;

stigma exserted beyond anthers.

Capsules

20–30 mm;

pedicel 0–5 mm.

10–35 mm;

pedicel 5–15 mm.

Seeds

brown or gray, 1.5–1.8 mm, scaly-echinate, crest 0.1 mm.

brown, 2 mm, minutely scaly, crest 0.5 mm, prominent.

2n

= 14.

Clarkia mildrediae

Clarkia arcuata

Phenology Flowering Apr–Jun.
Habitat Openings in woodlands and chaparral, serpentine soil.
Elevation 0–1600 m. (0–5200 ft.)
Distribution
from USDA
California
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Clarkia mildrediae is closely related to, and may be derived from, C. borealis. Clarkia mildrediae subsp. lutescens is probably the direct ancestor of C. stellata. Clarkia mildrediae differs from C. borealis in petal color and inflorescence habit and from C. stellata in flower size and position of the stigma. On the basis of morphology, chromosome number and pairing, C. mildrediae appears to be one of the parents of the tetraploid species C. rhomboidea.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Clarkia arcuata is primarily distributed in the foothills of the Sierra Nevada from Mariposa to Tehama counties, sparsely south to Kern County, and with one disjunct occurrence in Shasta County; it has also been reported from Napa County.

Clarkia arcuata is morphologically most similar to C. lassenensis, from which it differs in fruit characteristics. The two species have distinct areas of distribution and attempts to produce hybrids have not been successful.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals 12–18 mm wide; anthers magenta, fresh pollen blue-gray.
subsp. mildrediae
1. Petals 7–16 mm wide; anthers yellow to orange-red, fresh pollen bright yellow to tan.
subsp. lutescens
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Myxocarpa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Rhodanthos > subsect. Flexicaules
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. amoena, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. mildrediae subsp. lutescens, C. mildrediae subsp. mildrediae
Synonyms Phaeostoma mildrediae Oenothera arcuata, Godetia hispidula, O. hispidula
Name authority (A. Heller) H. Lewis & M. E. Lewis: Madroño 12: 34. (1953) (Kellogg) A. Nelson & J. F. Macbride: Bot. Gaz. 65: 62. (1918)
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