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Mildred's clarkia

arewell-to-spring, farewell-to-spring, farewell-to-spring clarkia, godetia, yellow clarkia

Stems

erect, to 100 cm, puberulent.

erect to decumbent, 20–200 cm, puberulent.

Leaves

petiole 15–40 mm;

blade elliptic to ovate, 3–6 cm.

petiole to 10 mm;

blade linear to lanceolate, 1–6 cm.

Inflorescences

open racemes, axis recurved in bud, straight only 1–3 nodes distal to open flowers;

buds pendent, tip acute.

open or dense spikes or racemes, axis straight;

buds erect.

Flowers

floral tube 2–3 mm;

sepals usually reflexed individually;

corolla rotate, petals reddish purple, often darker flecked or spotted, narrowly obdeltate to suborbiculate, unlobed, 11–25 × 7–18 mm, length 1.4–1.6 times width;

stamens 8, subequal, subtended by ciliate scales, anthers magenta or orange-red to yellow, pollen blue-gray or bright yellow to tan;

ovary shallowly 4-grooved, puberulent;

stigma exserted beyond anthers.

floral tube 3–10 mm;

sepals usually reflexed together to one side, or rarely in pairs or singly;

corolla bowl-shaped, petals pale pink to lavender, usually with red spot or mark near middle, obovate to fan-shaped, 15–60 mm, not lobed, apex sometimes shallowly notched or erose;

stamens 8, in 2 subequal sets;

ovary cylindrical and 4-grooved or fusiform and 8-grooved, puberulent;

stigma exserted or not beyond anthers.

Capsules

20–30 mm;

pedicel 0–5 mm.

15–40 mm, sometimes broader distally;

pedicel 0–13 mm.

Seeds

brown or gray, 1.5–1.8 mm, scaly-echinate, crest 0.1 mm.

brown to grayish brown, 1–1.5 mm, scaly, crest 0.1 mm.

Clarkia mildrediae

Clarkia amoena

Distribution
from USDA
California
[WildflowerSearch map]
[BONAP county map]
from USDA
w North America
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Clarkia mildrediae is closely related to, and may be derived from, C. borealis. Clarkia mildrediae subsp. lutescens is probably the direct ancestor of C. stellata. Clarkia mildrediae differs from C. borealis in petal color and inflorescence habit and from C. stellata in flower size and position of the stigma. On the basis of morphology, chromosome number and pairing, C. mildrediae appears to be one of the parents of the tetraploid species C. rhomboidea.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 5 (5 in the flora).

Clarkia amoena is closely related to C. rubicunda, which differs morphologically mainly in the color pattern of the petals. Petals of C. amoena have a conspicuous red spot or group of small red spots or marks near the middle, whereas those of C. rubicunda have a red area at the base and are not spotted near the middle. The areas of distribution of the two species barely overlap in California just north of San Francisco Bay, and C. rubicunda could be considered a southern geographical race or subspecies of C. amoena were it not that their readily formed hybrids are sterile due to chromosomal rearrangement. Clarkia amoena is one of the parent species of the allotetraploid C. gracilis. Intermediates between subspecies are frequent.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals 12–18 mm wide; anthers magenta, fresh pollen blue-gray.
subsp. mildrediae
1. Petals 7–16 mm wide; anthers yellow to orange-red, fresh pollen bright yellow to tan.
subsp. lutescens
1. Stigmas not exserted beyond anthers; petals less than 20 mm; coastal British Columbia, Oregon, Washington.
subsp. caurina
1. Stigmas exserted beyond anthers; petals 15–60 mm.
→ 2
2. Ovaries 8-grooved, broadly fusiform, 8–12 mm wide.
subsp. whitneyi
2. Ovaries 4-grooved, cylindrical to subclavate, 2–6 mm wide.
→ 3
3. Inflorescences open, internodes longer than subtending flowers, bracts sublinear.
subsp. huntiana
3. Inflorescences congested, internodes shorter than subtending flowers, bracts narrowly lanceolate or wider.
→ 4
4. Stems decumbent to suberect, to 100 cm; petals 20–35 mm, usually with red spot mid blade; coastal California.
subsp. amoena
4. Stems erect, to 200 cm; petals 30–40 mm, usually without red spot or with very small spot or streak mid blade; inland Oregon, Washington.
subsp. lindleyi
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Myxocarpa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Rhodanthos > subsect. Primigenia
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. mildrediae subsp. lutescens, C. mildrediae subsp. mildrediae
C. amoena subsp. amoena, C. amoena subsp. caurina, C. amoena subsp. huntiana, C. amoena subsp. lindleyi, C. amoena subsp. whitneyi
Synonyms Phaeostoma mildrediae Oenothera amoena, Godetia amoena
Name authority (A. Heller) H. Lewis & M. E. Lewis: Madroño 12: 34. (1953) (Lehmann) A. Nelson & J. F. Macbride: Bot. Gaz. 65: 62. (1918)
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