Clarkia mildrediae |
Clarkia affinis |
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Mildred's clarkia |
chaparral clarkia, chaparral clarkia or fairyfan, chaparral fairyfan |
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Stems | erect, to 100 cm, puberulent. |
erect, to 80 cm, puberulent. |
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Leaves | petiole 15–40 mm; blade elliptic to ovate, 3–6 cm. |
petiole 0–3 mm; blade linear to narrowly lanceolate, 1.5–7 cm. |
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Inflorescences | open racemes, axis recurved in bud, straight only 1–3 nodes distal to open flowers; buds pendent, tip acute. |
dense spikes, axis straight; buds erect. |
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Flowers | floral tube 2–3 mm; sepals usually reflexed individually; corolla rotate, petals reddish purple, often darker flecked or spotted, narrowly obdeltate to suborbiculate, unlobed, 11–25 × 7–18 mm, length 1.4–1.6 times width; stamens 8, subequal, subtended by ciliate scales, anthers magenta or orange-red to yellow, pollen blue-gray or bright yellow to tan; ovary shallowly 4-grooved, puberulent; stigma exserted beyond anthers. |
floral tube1.5–4 mm; sepals reflexed together to 1 side; corolla bowl-shaped, petals 5–15 mm; stamens 8, subequal; ovary cylindrical, 8-grooved, length at least 10 times width; stigma not exserted beyond anthers. |
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Capsules | 20–30 mm; pedicel 0–5 mm. |
15–30 mm, beak 3–7 mm; pedicel 0–5 mm. |
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Seeds | brown or gray, 1.5–1.8 mm, scaly-echinate, crest 0.1 mm. |
brown or gray, 1–1.5 mm, scaly, crest 0.1 mm. |
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2n | = 52. |
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Clarkia mildrediae |
Clarkia affinis |
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Phenology | Flowering Apr–Jun. | |||||
Habitat | Openings in woodlands and chaparral. | |||||
Elevation | 0–500 m. (0–1600 ft.) | |||||
Distribution |
California
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CA
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Discussion | Subspecies 2 (2 in the flora). Clarkia mildrediae is closely related to, and may be derived from, C. borealis. Clarkia mildrediae subsp. lutescens is probably the direct ancestor of C. stellata. Clarkia mildrediae differs from C. borealis in petal color and inflorescence habit and from C. stellata in flower size and position of the stigma. On the basis of morphology, chromosome number and pairing, C. mildrediae appears to be one of the parents of the tetraploid species C. rhomboidea. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Clarkia affinis is known primarily from west-central California and the North Coast Ranges, and more scattered in the Sierra Nevada Foothills and Western Transverse Ranges. Clarkia affinis is a hexaploid most closely related to C. purpurea; both have 2n = 52. Chromosome pairing in hybrids between them, as well as morphology, suggest that they have a tetraploid (2n = 34) genome in common. The two species are most readily distinguished by their immature capsules, which in C. affinis are slender, at least ten times longer than wide, beaked, and shallowly grooved, whereas those of C. purpurea are stout, not more than eight times longer than wide, not prominently beaked, and deeply grooved; the sepals of the former are generally reflexed together in fours whereas those of the latter are reflexed individually or in twos. Based on morphology and molecular data, the diploid genome probably came from C. cylindrica or a related species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 10. | FNA vol. 10. | ||||
Parent taxa | Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Myxocarpa | Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Biortis | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Phaeostoma mildrediae | |||||
Name authority | (A. Heller) H. Lewis & M. E. Lewis: Madroño 12: 34. (1953) | H. Lewis & M. E. Lewis: Madroño 12: 34. (1953) | ||||
Web links |