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Lewis' clarkia

Vine Hill clarkia

Stems

erect, to 50 cm, puberulent to glabrate.

erect, to 60 cm, glabrous or sparsely puberulent.

Leaves

petiole to 7 mm;

blade narrowly lanceolate to lanceolate, 2–5 cm.

petiole 0–2 mm;

blade lanceolate, 2–2.5 cm.

Inflorescences

open racemes, axis recurved at tip in bud;

buds pendent.

dense racemes, axis straight;

buds erect.

Flowers

floral tube 1.5–4 mm, with ring of hairs at distal margin inside;

sepals reflexed together to 1 side;

corolla bowl-shaped, petals pinkish lavender shading white near middle, base purplish red or with red line, sometimes reddish purple-flecked, 10–30 mm;

stamens 8, unequal, width of all filaments subequal or inner slightly thinner, outer anthers lavender, inner smaller, paler.

floral tube 10–15 mm, conspicuously veined, lavender striate within;

sepals reflexed individually;

corolla bowl-shaped, petals lavender shading to white proximally, with large, wedge-shaped purplish red spot near apex, 20–25 mm;

stamens 8, subequal;

ovary 8-grooved, longer than adjacent internode;

stigma exserted beyond anthers.

Capsules

15–70 mm, beak 0–3 mm.

10–15 mm.

Seeds

brown, 1 mm, scaly to puberulent, crest inconspicuous.

brown or gray, 2 mm, scaly, crest 0.2 mm.

2n

= 18.

= 16.

Clarkia lewisii

Clarkia imbricata

Phenology Flowering Jun. Flowering Jun–Jul.
Habitat Coastal scrub, woodlands, chaparral. Clearings, roadsides, chaparral.
Elevation 0–300 m. (0–1000 ft.) 50 m. (200 ft.)
Distribution
from FNA
CA
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Clarkia lewisii is known primarily from Monterey County, sparsely in San Benito County, barely reaching Santa Clara County, and is listed as rare by the California Native Plant Society. It is most closely related and morphologically similar to C. cylindrica, from which it can be distinguished by having all filaments about equally wide and a ring of hairs at the rim of the floral tube; outer filaments of C. cylindrica are two times as wide as the inner ones, and the ring of hairs is within the tube below the rim. Clarkia lewisii is also closely related to C. rostrata, from which it differs conspicuously by having a much shorter capsule beak.

Clarkia lewisii is a new name applied to the species known until 1978 as C. bottae, following examination and reinterpretation of the type of Godetia bottae Spach by P. H. Raven and D. R. Parnell (1978). They determined that the type specimens of G. bottae actually referred to the species then known as C. deflexa (Jepson) H. Lewis & M. E. Lewis, and reapplied the name C. bottae to that species in sect. Fibula.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Clarkia imbricata, known from only one small area of Sonoma County, is designated as rare by the California Native Plant Society, and is in the Center for Plant Conservation’s National Collection of Endangered Plants; it is a relict of a lineage with 2n = 16 that presumably contributed a genome to the tetraploid species C. davyi and the South American C. tenella.

Morphologically, Clarkia imbricata is most similar to C. speciosa, C. williamsonii, and some populations of C. purpurea. Clarkia imbricata can be distinguished from C. speciosa by the color pattern of the petals and from C. williamsonii and populations of C. purpurea with similar flower size and color pattern by its broader, ascending, overlapping leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Phaeostoma > subsect. Sympherica Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Godetia
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Name authority P. H. Raven & D. R. Parnell: Ann. Missouri Bot. Gard. 64: 642. (1978) H. Lewis & M. E. Lewis: Madroño 12: 38. (1953)
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