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Vine Hill clarkia

Mildred's clarkia

Stems

erect, to 60 cm, glabrous or sparsely puberulent.

erect, to 100 cm, puberulent.

Leaves

petiole 0–2 mm;

blade lanceolate, 2–2.5 cm.

petiole 15–40 mm;

blade elliptic to ovate, 3–6 cm.

Inflorescences

dense racemes, axis straight;

buds erect.

open racemes, axis recurved in bud, straight only 1–3 nodes distal to open flowers;

buds pendent, tip acute.

Flowers

floral tube 10–15 mm, conspicuously veined, lavender striate within;

sepals reflexed individually;

corolla bowl-shaped, petals lavender shading to white proximally, with large, wedge-shaped purplish red spot near apex, 20–25 mm;

stamens 8, subequal;

ovary 8-grooved, longer than adjacent internode;

stigma exserted beyond anthers.

floral tube 2–3 mm;

sepals usually reflexed individually;

corolla rotate, petals reddish purple, often darker flecked or spotted, narrowly obdeltate to suborbiculate, unlobed, 11–25 × 7–18 mm, length 1.4–1.6 times width;

stamens 8, subequal, subtended by ciliate scales, anthers magenta or orange-red to yellow, pollen blue-gray or bright yellow to tan;

ovary shallowly 4-grooved, puberulent;

stigma exserted beyond anthers.

Capsules

10–15 mm.

20–30 mm;

pedicel 0–5 mm.

Seeds

brown or gray, 2 mm, scaly, crest 0.2 mm.

brown or gray, 1.5–1.8 mm, scaly-echinate, crest 0.1 mm.

2n

= 16.

Clarkia imbricata

Clarkia mildrediae

Phenology Flowering Jun–Jul.
Habitat Clearings, roadsides, chaparral.
Elevation 50 m. (200 ft.)
Distribution
from FNA
CA
[BONAP county map]
from USDA
California
[WildflowerSearch map]
[BONAP county map]
Discussion

Clarkia imbricata, known from only one small area of Sonoma County, is designated as rare by the California Native Plant Society, and is in the Center for Plant Conservation’s National Collection of Endangered Plants; it is a relict of a lineage with 2n = 16 that presumably contributed a genome to the tetraploid species C. davyi and the South American C. tenella.

Morphologically, Clarkia imbricata is most similar to C. speciosa, C. williamsonii, and some populations of C. purpurea. Clarkia imbricata can be distinguished from C. speciosa by the color pattern of the petals and from C. williamsonii and populations of C. purpurea with similar flower size and color pattern by its broader, ascending, overlapping leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

Clarkia mildrediae is closely related to, and may be derived from, C. borealis. Clarkia mildrediae subsp. lutescens is probably the direct ancestor of C. stellata. Clarkia mildrediae differs from C. borealis in petal color and inflorescence habit and from C. stellata in flower size and position of the stigma. On the basis of morphology, chromosome number and pairing, C. mildrediae appears to be one of the parents of the tetraploid species C. rhomboidea.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals 12–18 mm wide; anthers magenta, fresh pollen blue-gray.
subsp. mildrediae
1. Petals 7–16 mm wide; anthers yellow to orange-red, fresh pollen bright yellow to tan.
subsp. lutescens
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Godetia Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Myxocarpa
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. mildrediae subsp. lutescens, C. mildrediae subsp. mildrediae
Synonyms Phaeostoma mildrediae
Name authority H. Lewis & M. E. Lewis: Madroño 12: 38. (1953) (A. Heller) H. Lewis & M. E. Lewis: Madroño 12: 34. (1953)
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