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canyon clarkia, willow-herb clarkia

graceful clarkia, slender clarkia, slender godetia

Stems

erect, 20–70 cm, sparsely puberulent.

erect, to 90 cm, glabrous or puberulent.

Leaves

petiole to 7 mm;

blade linear to narrowly lanceolate or oblanceolate, 1.5–2.5 cm.

petiole to 11 mm;

blade linear to narrowly lanceolate, 2–7 cm.

Inflorescences

open racemes, sometimes few-branched, axis recurved at tip in bud;

buds pendent.

racemes, axis recurved at tip in bud;

buds pendent.

Flowers

usually cleistogamous;

floral tube 1–3 mm;

sepals reflexed together to 1 side or in pairs;

corolla bowl-shaped, petals fading pink, obovate;

stamens 8, unequal, anthers white or cream, outer ones larger than inner.

floral tube 1.5–10 mm;

corolla bowl-shaped, petals pink to lavender or light purple, often lighter toward base, unspotted or with red spot near middle or base, obovate or obdeltate, 6–40 mm;

stamens 8, subequal;

ovary 4-grooved, puberulent;

stigma exserted or not beyond anthers.

Capsules

10–30 mm;

pedicel 5–11 mm.

25–50 mm, usually wider distally;

pedicel 0–10 mm.

Seeds

brown, 0.5–1 mm, scaly, crest inconspicuous.

brown or mottled with dark spots, 1.5–2 mm, scaly, crest 0.1 mm.

2n

= 18.

Clarkia epilobioides

Clarkia gracilis

Phenology Flowering Apr–May.
Habitat Shady sites, woodlands, chaparral.
Elevation 0–1000 m. (0–3300 ft.)
Distribution
from FNA
AZ; CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from USDA
w United States
[WildflowerSearch map]
[BONAP county map]
Discussion

Clarkia epilobioides is known from south-central Arizona in Gila, Maricopa, Pima, and Pinal counties, and widely in west-central and southwestern California and adjacent Baja California, Mexico. In California, it occurs from Contra Costa and San Mateo counties in the San Francisco Bay area to San Diego County in the south, including most of the Channel Islands.

Clarkia epilobioides is modally self-pollinating, and up to half of its flowers do not open, yet set a full complement of seeds (H. Lewis and M. E. Lewis 1955). However, outcrossing does occur, and C. epilobioides is one of the parents of the tetraploid species C. similis, from which it differs by having white, unflecked petals; it is also one of the parents of the tetraploid C. delicata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 4 (4 in the flora).

Clarkia gracilis is a tetraploid derived from C. amoena and an unknown, presumably extinct, species related to C. arcuata and C. lassenensis. Clarkia gracilis is readily distinguished from C. amoena by the axis of the inflorescence in bud, which is not recurved in C. amoena; it is distinguished from C. arcuata and C. lassenensis by its four rather than eight-grooved ovary.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stigmas not exserted beyond anthers; petals 6–23 mm.
subsp. gracilis
1. Stigmas exserted beyond anthers; petals20–40 mm.
→ 2
2. Petals with red spot near mid blade, base of blades white.
subsp. sonomensis
2. Petals unspotted, base of blades red or bright red.
→ 3
3. Petals 20–30 mm.
subsp. tracyi
3. Petals 30–40 mm.
subsp. albicaulis
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Phaeostoma > subsect. Micranthae Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Rhodanthos > subsect. Jugales
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. gracilis subsp. albicaulis, C. gracilis subsp. gracilis, C. gracilis subsp. sonomensis, C. gracilis subsp. tracyi
Synonyms Oenothera epilobioides, Godetia epilobioides Godetia gracilis, G. amoena var. gracilis
Name authority (Nuttall ex Torrey & A. Gray) A. Nelson & J. F. Macbride: Bot. Gaz. 65: 60. (1918) (Piper) A. Nelson & J. F. Macbride: Bot. Gaz. 65: 63. (1918)
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