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canyon clarkia, willow-herb clarkia

speckled clarkia, speckled fairyfan

Stems

erect, 20–70 cm, sparsely puberulent.

erect, to 60 cm, puberulent or glabrous.

Leaves

petiole to 7 mm;

blade linear to narrowly lanceolate or oblanceolate, 1.5–2.5 cm.

petiole to 5 mm;

blade linear to narrowly lanceolate, 1–6 cm.

Inflorescences

open racemes, sometimes few-branched, axis recurved at tip in bud;

buds pendent.

open racemes, axis recurved at tip in bud;

buds pendent.

Flowers

usually cleistogamous;

floral tube 1–3 mm;

sepals reflexed together to 1 side or in pairs;

corolla bowl-shaped, petals fading pink, obovate;

stamens 8, unequal, anthers white or cream, outer ones larger than inner.

floral tube 2–7 mm, with ring of hairs proximal to distal margin inside;

sepals reflexed together to 1 side;

corolla bowl-shaped, petals purple to pinkish lavender shading white near middle, often reddish purple-flecked, base bright purplish red, 10–35 mm;

stamens 8, unequal, width of outer filaments about 2 times inner, outer anthers lavender, inner smaller, paler.

Capsules

10–30 mm;

pedicel 5–11 mm.

20–50 mm, beak 3–5 mm.

Seeds

brown, 0.5–1 mm, scaly, crest inconspicuous.

brown, 1–1.5 mm, minutely scaly to puberulent, crest 0.1 mm.

2n

= 18.

Clarkia epilobioides

Clarkia cylindrica

Phenology Flowering Apr–May.
Habitat Shady sites, woodlands, chaparral.
Elevation 0–1000 m. (0–3300 ft.)
Distribution
from FNA
AZ; CA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
from USDA
California
[WildflowerSearch map]
[BONAP county map]
Discussion

Clarkia epilobioides is known from south-central Arizona in Gila, Maricopa, Pima, and Pinal counties, and widely in west-central and southwestern California and adjacent Baja California, Mexico. In California, it occurs from Contra Costa and San Mateo counties in the San Francisco Bay area to San Diego County in the south, including most of the Channel Islands.

Clarkia epilobioides is modally self-pollinating, and up to half of its flowers do not open, yet set a full complement of seeds (H. Lewis and M. E. Lewis 1955). However, outcrossing does occur, and C. epilobioides is one of the parents of the tetraploid species C. similis, from which it differs by having white, unflecked petals; it is also one of the parents of the tetraploid C. delicata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

As defined by Davis, the subspecies of Clarkia cylindrica have distinct but partly overlapping geographical ranges; subsp. cylindrica mainly in the South Coast and Transverse Ranges to the Tehachapi Mountain area, and subsp. clavicarpa mainly in the central and southern Sierra Nevada Foothills to the Tehachapi Mountain area. More recent collections suggest more substantial geographical overlap. Morphological variation correlates with geographical distribution, with the most consistent difference in ovary and capsule shape. According to Davis, the taxa are moderately interfertile, less so for more distantly separated individuals.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Ovaries and capsules cylindrical.
subsp. cylindrica
1. Ovaries and capsules subclavate, enlarged distally.
subsp. clavicarpa
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Phaeostoma > subsect. Micranthae Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Phaeostoma > subsect. Sympherica
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. cylindrica subsp. clavicarpa, C. cylindrica subsp. cylindrica
Synonyms Oenothera epilobioides, Godetia epilobioides Godetiabottae spach var. cylindrica
Name authority (Nuttall ex Torrey & A. Gray) A. Nelson & J. F. Macbride: Bot. Gaz. 65: 60. (1918) (Jepson) H. Lewis & M. E. Lewis: Madroño 12: 33. (1953) — (as cyclindrica)
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