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Dudley's clarkia

California gaura, heterogaura, mountain clarkia

Stems

erect, to 70 cm, puberulent.

erect, to 60 cm, glandular puberulent.

Leaves

petiole 3–10 mm;

blade narrowly lanceolate, 1.5–7 cm.

petiole 5–20 mm;

blade lanceolate to ovate, 2–8 cm.

Inflorescences

open racemes, axis recurved at tip in bud;

buds pendent.

open racemes or panicles, axis straight;

buds erect.

Flowers

floral tube 1–3 mm;

sepals reflexed together to 1 side;

corolla bowl-shaped, petals lavender-pink, usually white-streaked, often red-flecked, broadly fan-shaped, 10–30 mm, apex subentire to crenulate;

stamens 8, unequal, outer anthers lavender, inner smaller, paler.

sometimes autogamous;

floral tube 1–2 mm;

sepals reflexed together to 1 side;

corolla rotate, petals elliptic to obovate, tapered to claw;

stamens 8, unequal, anthers cream or light pink, inner ones smaller, sterile;

ovary subglobose, grooves obscure;

stigma not exserted beyond anthers.

Capsules

10–30 mm.

2–3 mm, indehiscent, nutlike;

sessile or pedicellate (to 2 mm).

Seeds

brown, 1 mm, minutely scaly to puberulent, crest inconspicuous.

1 or 2.

2n

= 18.

= 18.

Clarkia dudleyana

Clarkia heterandra

Phenology Flowering May–Jul. Flowering May–Jul.
Habitat Openings in woodlands, chaparral, yellow-pine forests, coastal sage. Shady sites, woodlands, yellow-pine forests.
Elevation 0–1500 m. (0–4900 ft.) 500–2000 m. (1600–6600 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Clarkia dudleyana is a rather widespread species in California, known primarily from the central and southern Sierra Nevada foothills, the Tehachapi Mountain area, the Transverse Ranges, and the Peninsular Ranges, ranging from Tuolumne to Riverside counties, sporadically in the north to Nevada County and in the south to San Diego County

Clarkia dudleyana is morphologically most similar to C. biloba and C. modesta, but molecular data suggest that the relationship is not close. On the basis of chloroplast DNA sequence, C. dudleyana and C. heterandra are closely related.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Clarkia heterandra occurs in southern Oregon and California, where it is known from the Klamath Ranges in Trinity County, widely in the Sierra Nevada and Tehachapi Mountain area from Nevada to Kern counties, and in the South Coast and Transverse Ranges from San Luis Obispo to Riverside and San Bernardino counties.

Prior to 1986, Clarkia heterandra was treated as the monotypic genus Heterogaura, thought to be possibly related to the genus Gaura (now a section of Oenothera) due to its similar indehiscent fruits. However, molecular data indicate that C. heterandra is clearly within Clarkia, most closely related to C. dudleyana and C. unguiculata, which suggests a possible hybrid origin at the diploid level (K. J. Sytsma and L. D. Gottlieb 1986, 1986b; W. L. Wagner et al. 2007). Clarkia dudleyana and C. unguiculata produce spontaneous hybrids when grown adjacent to each other if they come from allopatric populations.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Phaeostoma > subsect. Lautiflorae Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Phaeostoma > subsect. Heterogaura
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Synonyms Godetia dudleyana Gaura heterandra, Heterogaura heterandra
Name authority (Abrams) J. F. Macbride: Contr. Gray Herb. 56: 54. (1918) (Torrey) H. Lewis & P. H. Raven: Madroño 39: 163. (1992)
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