Clarkia cylindrica
Onagraceae subfam. onagroideae
speckled clarkia, speckled fairyfan
erect, to 60 cm, puberulent or glabrous.
petiole to 5 mm;
blade linear to narrowly lanceolate, 1–6 cm.
stipules present or absent.
open racemes, axis recurved at tip in bud;
buds pendent.
floral tube 2–7 mm, with ring of hairs proximal to distal margin inside;
sepals reflexed together to 1 side;
corolla bowl-shaped, petals purple to pinkish lavender shading white near middle, often reddish purple-flecked, base bright purplish red, 10–35 mm;
stamens 8, unequal, width of outer filaments about 2 times inner, outer anthers lavender, inner smaller, paler.
floral tube present or, rarely, absent;
sepals 2 or 4 (very rarely 3), deciduous with floral tube, petals, and stamens;
petals yellow, white, pink, red, rarely in combination.
20–50 mm, beak 3–5 mm.
brown, 1–1.5 mm, minutely scaly to puberulent, crest 0.1 mm.
Clarkia cylindrica
Onagraceae subfam. onagroideae
Subspecies 2 (2 in the flora).
As defined by Davis, the subspecies of Clarkia cylindrica have distinct but partly overlapping geographical ranges; subsp. cylindrica mainly in the South Coast and Transverse Ranges to the Tehachapi Mountain area, and subsp. clavicarpa mainly in the central and southern Sierra Nevada Foothills to the Tehachapi Mountain area. More recent collections suggest more substantial geographical overlap. Morphological variation correlates with geographical distribution, with the most consistent difference in ovary and capsule shape. According to Davis, the taxa are moderately interfertile, less so for more distantly separated individuals.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera 21, species 582 (16 genera, 246 species in the flora).
Onagroideae encompass the main lineage of the family, after the early branching of Ludwigia (R. A. Levin et al. 2003, 2004). This large and diverse lineage is distinguished by the presence of a floral tube beyond the apex of the ovary; sepals deciduous with the floral tube, petals, and stamens; pollen shed in monads (or tetrads in Chylismia sect. Lignothera and all but one species of Epilobium); ovular vascular system exclusively transseptal (R. H. Eyde 1981); ovule archesporium multicellular (H. Tobe and P. H. Raven 1996); and change in base chromosome number from x = 8 in Ludwigia to x = 10 or x = 11 at the base of Onagroideae (Raven 1979; Levin et al. 2003). Molecular work (Levin et al. 2003, 2004) substantially supports the traditional tribal classification (P. A. Munz 1965; Raven 1979, 1988); tribes are recognized to delimit major branches within the phylogeny of Onagroideae, where the branches comprise strongly supported monophyletic groups of one or more genera.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Ovaries and capsules cylindrical. | subsp. cylindrica |
1. Ovaries and capsules subclavate, enlarged distally. | subsp. clavicarpa |
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