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speckled clarkia, speckled fairyfan

purple clarkia, winecup clarkia, winecup fairyfan

Stems

erect, to 60 cm, puberulent or glabrous.

erect or rarely decumbent, to 100 cm, glabrous and sometimes glaucous or sparsely to densely puberulent, sometimes mixed with longer, spreading hairs.

Leaves

petiole to 5 mm;

blade linear to narrowly lanceolate, 1–6 cm.

petiole 0–2 mm;

blade linear or narrowly lanceolate to elliptic or ovate, 1.5–7 cm.

Inflorescences

open racemes, axis recurved at tip in bud;

buds pendent.

open or dense racemes, axis straight;

buds erect.

Flowers

floral tube 2–7 mm, with ring of hairs proximal to distal margin inside;

sepals reflexed together to 1 side;

corolla bowl-shaped, petals purple to pinkish lavender shading white near middle, often reddish purple-flecked, base bright purplish red, 10–35 mm;

stamens 8, unequal, width of outer filaments about 2 times inner, outer anthers lavender, inner smaller, paler.

floral tube 2–10 mm;

sepals reflexed individually or in pairs;

corolla bowl-shaped, petals lavender to purple, purplish red, or dark wine-red, often with red or purple spot near middle, tip, or base, 9–25 mm;

stamens 8, subequal;

ovary 8-grooved, length less than 8 times width;

stigma as long as or exserted beyond anthers.

Capsules

20–50 mm, beak 3–5 mm.

10–30 mm, beak 0–2 mm.

Seeds

brown, 1–1.5 mm, minutely scaly to puberulent, crest 0.1 mm.

brown or gray, 1–2 mm, scaly, crest 0.2 mm.

Clarkia cylindrica

Clarkia purpurea

Distribution
from USDA
California
[WildflowerSearch map]
[BONAP county map]
from USDA
w North America; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

As defined by Davis, the subspecies of Clarkia cylindrica have distinct but partly overlapping geographical ranges; subsp. cylindrica mainly in the South Coast and Transverse Ranges to the Tehachapi Mountain area, and subsp. clavicarpa mainly in the central and southern Sierra Nevada Foothills to the Tehachapi Mountain area. More recent collections suggest more substantial geographical overlap. Morphological variation correlates with geographical distribution, with the most consistent difference in ovary and capsule shape. According to Davis, the taxa are moderately interfertile, less so for more distantly separated individuals.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 3 (3 in the flora).

Clarkia purpurea consists of a diverse assemblage of hexaploid populations and is almost certainly derived from multiple origins followed by hybridization and, perhaps, backcrossing to parental species. Three morphological forms are recognized as subspecies; intergrades are frequent.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Ovaries and capsules cylindrical.
subsp. cylindrica
1. Ovaries and capsules subclavate, enlarged distally.
subsp. clavicarpa
1. Leaf blades broadly lanceolate to elliptic or ovate; inflorescences dense racemes.
subsp. purpurea
1. Leaf blades linear to lanceolate or narrowly lanceolate; inflorescences usually open racemes.
→ 2
2. Petals 15–25 mm; stigmas exserted beyond anthers.
subsp. viminea
2. Petals 9–14 mm; stigmas not exserted beyond anthers.
subsp. quadrivulnera
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Phaeostoma > subsect. Sympherica Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Godetia
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. cylindrica subsp. clavicarpa, C. cylindrica subsp. cylindrica
C. purpurea subsp. purpurea, C. purpurea subsp. quadrivulnera, C. purpurea subsp. viminea
Synonyms Godetiabottae spach var. cylindrica Oenothera purpurea, Godetia purpurea
Name authority (Jepson) H. Lewis & M. E. Lewis: Madroño 12: 33. (1953) — (as cyclindrica) (Curtis) A. Nelson & J. F. Macbride: Bot. Gaz. 65: 64. (1918)
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