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speckled clarkia, speckled fairyfan

chaparral clarkia, chaparral clarkia or fairyfan, chaparral fairyfan

Stems

erect, to 60 cm, puberulent or glabrous.

erect, to 80 cm, puberulent.

Leaves

petiole to 5 mm;

blade linear to narrowly lanceolate, 1–6 cm.

petiole 0–3 mm;

blade linear to narrowly lanceolate, 1.5–7 cm.

Inflorescences

open racemes, axis recurved at tip in bud;

buds pendent.

dense spikes, axis straight;

buds erect.

Flowers

floral tube 2–7 mm, with ring of hairs proximal to distal margin inside;

sepals reflexed together to 1 side;

corolla bowl-shaped, petals purple to pinkish lavender shading white near middle, often reddish purple-flecked, base bright purplish red, 10–35 mm;

stamens 8, unequal, width of outer filaments about 2 times inner, outer anthers lavender, inner smaller, paler.

floral tube1.5–4 mm;

sepals reflexed together to 1 side;

corolla bowl-shaped, petals 5–15 mm;

stamens 8, subequal;

ovary cylindrical, 8-grooved, length at least 10 times width;

stigma not exserted beyond anthers.

Capsules

20–50 mm, beak 3–5 mm.

15–30 mm, beak 3–7 mm;

pedicel 0–5 mm.

Seeds

brown, 1–1.5 mm, minutely scaly to puberulent, crest 0.1 mm.

brown or gray, 1–1.5 mm, scaly, crest 0.1 mm.

2n

= 52.

Clarkia cylindrica

Clarkia affinis

Phenology Flowering Apr–Jun.
Habitat Openings in woodlands and chaparral.
Elevation 0–500 m. (0–1600 ft.)
Distribution
from USDA
California
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

As defined by Davis, the subspecies of Clarkia cylindrica have distinct but partly overlapping geographical ranges; subsp. cylindrica mainly in the South Coast and Transverse Ranges to the Tehachapi Mountain area, and subsp. clavicarpa mainly in the central and southern Sierra Nevada Foothills to the Tehachapi Mountain area. More recent collections suggest more substantial geographical overlap. Morphological variation correlates with geographical distribution, with the most consistent difference in ovary and capsule shape. According to Davis, the taxa are moderately interfertile, less so for more distantly separated individuals.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Clarkia affinis is known primarily from west-central California and the North Coast Ranges, and more scattered in the Sierra Nevada Foothills and Western Transverse Ranges.

Clarkia affinis is a hexaploid most closely related to C. purpurea; both have 2n = 52. Chromosome pairing in hybrids between them, as well as morphology, suggest that they have a tetraploid (2n = 34) genome in common. The two species are most readily distinguished by their immature capsules, which in C. affinis are slender, at least ten times longer than wide, beaked, and shallowly grooved, whereas those of C. purpurea are stout, not more than eight times longer than wide, not prominently beaked, and deeply grooved; the sepals of the former are generally reflexed together in fours whereas those of the latter are reflexed individually or in twos. Based on morphology and molecular data, the diploid genome probably came from C. cylindrica or a related species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Ovaries and capsules cylindrical.
subsp. cylindrica
1. Ovaries and capsules subclavate, enlarged distally.
subsp. clavicarpa
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Phaeostoma > subsect. Sympherica Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Biortis
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. cylindrica subsp. clavicarpa, C. cylindrica subsp. cylindrica
Synonyms Godetiabottae spach var. cylindrica
Name authority (Jepson) H. Lewis & M. E. Lewis: Madroño 12: 33. (1953) — (as cyclindrica) H. Lewis & M. E. Lewis: Madroño 12: 34. (1953)
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