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botta's clarkia, punch bowl godetia

gunsight clarkia, Xantus' clarkia

Stems

erect, to 100 cm, glabrous, glaucous, rarely puberulent.

erect, to 80 cm, glabrous, glaucous.

Leaves

petiole to 5 mm;

blade narrowly lanceolate to lanceolate, 3–10 cm.

petiole 0–2 mm;

blade linear to lanceolate, 2–6 cm.

Inflorescences

open racemes, axis straight in bud;

buds pendent.

open racemes, axis straight;

buds pendent.

Flowers

floral tube 2–3 mm;

sepals reflexed together to 1 side;

corolla bowl-shaped, petals pale lavender to pinkish lavender, often white toward base, usually red-flecked, unlobed, 15–30 mm;

stamens 8, unequal, outer anthers lavender, inner smaller, paler;

ovary obscurely 4-grooved;

stigma exserted beyond anthers.

floral tube 2–5 mm;

sepals reflexed together to 1 side;

corolla rotate, petals lavender to reddish purple, lavender-pink, or white, often with white-surrounded dark reddish purple spot distally, clawed, 2-lobed, with slender central tooth, 1–3 mm, 6–20 mm;

stamens 8, in 2 unequal sets, outer anthers lavender to purple, inner smaller, paler;

ovary 8-grooved;

stigma exserted or not beyond anthers.

Capsules

30–40 mm;

pedicel 0–30 mm.

15–25 mm;

pedicel 0–5 mm.

Seeds

brown or gray, 1.2–1.8 mm, ± papillose, crest 0.2 mm.

brown, 1.3–1.5 mm, tuberculate, crest minute.

2n

= 18.

Clarkia bottae

Clarkia xantiana

Phenology Flowering Apr–Jul.
Habitat Chaparral, woodlands, coastal scrub.
Elevation 0–1000 m. (0–3300 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from USDA
California
[WildflowerSearch map]
[BONAP county map]
Discussion

Clarkia bottae is known from the Outer South Coast Ranges from Monterey and San Benito to Santa Barbara counties, throughout southwestern California, and more sporadically in the southern Sierra Nevada in Fresno and Kern counties. Reports from Del Norte, Madera, and Napa counties have not been verified.

Clarkia bottae is most closely related to C. jolonensis and may be closely related to C. xantiana; all three have 2n = 18, and spontaneous hybrids were formed when C. bottae was grown adjacent to a natural population of C. xantiana.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

Originally placed in the group now delimited as subsect. Phaeostoma by H. Lewis and M. E. Lewis (1955), Clarkia xantiana was treated as a monotypic subsect. Xantianae within sect. Phaeostoma by K. E. Holsinger (1985), based mainly on its unusual 2-lobed petals with a tooth in the sinus. Molecular data (R. A. Levin et al. 2004) placed C. xantiana close to C. bottae, and both of them close to but not within sect. Phaeostoma. Both species share the chromosome number 2n = 18 with C. jolonensis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals 12–20 mm; stigmas exserted beyond anthers.
subsp. xantiana
1. Petals 6–12 mm; stigmas not exserted beyond anthers.
subsp. parviflora
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Fibula Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Fibula
Sibling taxa
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii
Subordinate taxa
C. xantiana subsp. parviflora, C. xantiana subsp. xantiana
Synonyms Godetia bottae, C. deflexa, G. bottae var. deflexa, G. deflexa
Name authority (Spach) H. Lewis & M. E. Lewis: Madroño 12: 33. (1953) A. Gray: Proc. Boston Soc. Nat. Hist. 7: 146. (1859)
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