Clarkia amoena
Onagraceae tribe Onagreae
arewell-to-spring, farewell-to-spring, farewell-to-spring clarkia, godetia, yellow clarkia
erect to decumbent, 20–200 cm, puberulent.
petiole to 10 mm;
blade linear to lanceolate, 1–6 cm.
alternate or basal;
stipules absent.
open or dense spikes or racemes, axis straight;
buds erect.
floral tube 3–10 mm;
sepals usually reflexed together to one side, or rarely in pairs or singly;
corolla bowl-shaped, petals pale pink to lavender, usually with red spot or mark near middle, obovate to fan-shaped, 15–60 mm, not lobed, apex sometimes shallowly notched or erose;
stamens 8, in 2 subequal sets;
ovary cylindrical and 4-grooved or fusiform and 8-grooved, puberulent;
stigma exserted or not beyond anthers.
usually actinomorphic, rarely slightly zygomorphic (in Oenothera), (3 or)4-merous;
stamens 2 times as many, or rarely as many, as sepals;
pollen usually shed in monads, rarely tetrads (Chylismia sect. Lignothera).
a dry capsule, usually dehiscent, sometimes indehiscent.
15–40 mm, sometimes broader distally;
pedicel 0–13 mm.
brown to grayish brown, 1–1.5 mm, scaly, crest 0.1 mm.
few to numerous, without hairs or wings, [very rarely with asymmetrical dry wing (Xylonagra)], or with dry (Oenothera), erose or smooth wing, or with thick, papillate wings (Chylismiella).
Clarkia amoena
Onagraceae tribe Onagreae
Subspecies 5 (5 in the flora).
Clarkia amoena is closely related to C. rubicunda, which differs morphologically mainly in the color pattern of the petals. Petals of C. amoena have a conspicuous red spot or group of small red spots or marks near the middle, whereas those of C. rubicunda have a red area at the base and are not spotted near the middle. The areas of distribution of the two species barely overlap in California just north of San Francisco Bay, and C. rubicunda could be considered a southern geographical race or subspecies of C. amoena were it not that their readily formed hybrids are sterile due to chromosomal rearrangement. Clarkia amoena is one of the parent species of the allotetraploid C. gracilis. Intermediates between subspecies are frequent.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Genera 13, species 265 (12 genera, 199 species in the flora).
Onagreae account for more than half the total genera in Onagraceae and diversified from a center in southwestern North America (L. Katinas et al. 2004). Delimitation of the tribe by W. L. Wagner et al. (2007) differs from previous ones by the exclusion of Gongylocarpus, now in its own tribe, by the segregation of eight genera (Camissoniopsis, Chylismia, Chylismiella, Eremothera, Eulobus, Neoholmgrenia, Taraxia, and Tetrapteron) from Camissonia, and by the inclusion of three previously separate genera (Calylophus, Gaura, and Stenosiphon) in Oenothera. Within the branch of the family that lacks stipules (Gongylocarpeae, Epilobieae, and Onagreae), the last two tribes form a clade that has very strong molecular support (R. A. Levin et al. 2003, 2004), but no obvious morphological synapomorphy. The clade may be defined by a cytogenetic change from the base chromosome number of x = 11 found in Circaeeae, Gongylocarpeae, and Lopezieae, to x = 18 in Epilobieae, and x = 7 in Onagreae; however, these changes could also have occurred independently. Other than the new chromosome number x = 7, the only apparent morphological synapomorphy for Onagreae alone is pollen with prominent apertural protrusions (J. Praglowski et al. 1987, 1989), a character state also found in Circaeeae (Praglowski et al. 1994). The monophyly of Onagreae has moderate (Levin et al. 2004) to strong support (V. S. Ford and L. D. Gottlieb 2007).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Stigmas not exserted beyond anthers; petals less than 20 mm; coastal British Columbia, Oregon, Washington. | subsp. caurina |
1. Stigmas exserted beyond anthers; petals 15–60 mm. | → 2 |
2. Ovaries 8-grooved, broadly fusiform, 8–12 mm wide. | subsp. whitneyi |
2. Ovaries 4-grooved, cylindrical to subclavate, 2–6 mm wide. | → 3 |
3. Inflorescences open, internodes longer than subtending flowers, bracts sublinear. | subsp. huntiana |
3. Inflorescences congested, internodes shorter than subtending flowers, bracts narrowly lanceolate or wider. | → 4 |
4. Stems decumbent to suberect, to 100 cm; petals 20–35 mm, usually with red spot mid blade; coastal California. | subsp. amoena |
4. Stems erect, to 200 cm; petals 30–40 mm, usually without red spot or with very small spot or streak mid blade; inland Oregon, Washington. | subsp. lindleyi |
- Local floras:
BC, 
CA, 
OR, 
WA - Local Web sites: CalFlora, CalPhotos,
Flora NW, 
PNW Herbaria, Turner Photog. 
WildflowerSearch
iNaturalist (observations)
USDA Plants Database- LBJ Wildflower Center
- SEINet
- Plants of the World Online
- Encyclopedia of Life
- Wikipedia
- Google Image Search
