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chaparral clarkia, chaparral clarkia or fairyfan, chaparral fairyfan

purple clarkia, winecup clarkia, winecup fairyfan

Stems

erect, to 80 cm, puberulent.

erect or rarely decumbent, to 100 cm, glabrous and sometimes glaucous or sparsely to densely puberulent, sometimes mixed with longer, spreading hairs.

Leaves

petiole 0–3 mm;

blade linear to narrowly lanceolate, 1.5–7 cm.

petiole 0–2 mm;

blade linear or narrowly lanceolate to elliptic or ovate, 1.5–7 cm.

Inflorescences

dense spikes, axis straight;

buds erect.

open or dense racemes, axis straight;

buds erect.

Flowers

floral tube1.5–4 mm;

sepals reflexed together to 1 side;

corolla bowl-shaped, petals 5–15 mm;

stamens 8, subequal;

ovary cylindrical, 8-grooved, length at least 10 times width;

stigma not exserted beyond anthers.

floral tube 2–10 mm;

sepals reflexed individually or in pairs;

corolla bowl-shaped, petals lavender to purple, purplish red, or dark wine-red, often with red or purple spot near middle, tip, or base, 9–25 mm;

stamens 8, subequal;

ovary 8-grooved, length less than 8 times width;

stigma as long as or exserted beyond anthers.

Capsules

15–30 mm, beak 3–7 mm;

pedicel 0–5 mm.

10–30 mm, beak 0–2 mm.

Seeds

brown or gray, 1–1.5 mm, scaly, crest 0.1 mm.

brown or gray, 1–2 mm, scaly, crest 0.2 mm.

2n

= 52.

Clarkia affinis

Clarkia purpurea

Phenology Flowering Apr–Jun.
Habitat Openings in woodlands and chaparral.
Elevation 0–500 m. (0–1600 ft.)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from USDA
w North America; nw Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Clarkia affinis is known primarily from west-central California and the North Coast Ranges, and more scattered in the Sierra Nevada Foothills and Western Transverse Ranges.

Clarkia affinis is a hexaploid most closely related to C. purpurea; both have 2n = 52. Chromosome pairing in hybrids between them, as well as morphology, suggest that they have a tetraploid (2n = 34) genome in common. The two species are most readily distinguished by their immature capsules, which in C. affinis are slender, at least ten times longer than wide, beaked, and shallowly grooved, whereas those of C. purpurea are stout, not more than eight times longer than wide, not prominently beaked, and deeply grooved; the sepals of the former are generally reflexed together in fours whereas those of the latter are reflexed individually or in twos. Based on morphology and molecular data, the diploid genome probably came from C. cylindrica or a related species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 3 (3 in the flora).

Clarkia purpurea consists of a diverse assemblage of hexaploid populations and is almost certainly derived from multiple origins followed by hybridization and, perhaps, backcrossing to parental species. Three morphological forms are recognized as subspecies; intergrades are frequent.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades broadly lanceolate to elliptic or ovate; inflorescences dense racemes.
subsp. purpurea
1. Leaf blades linear to lanceolate or narrowly lanceolate; inflorescences usually open racemes.
→ 2
2. Petals 15–25 mm; stigmas exserted beyond anthers.
subsp. viminea
2. Petals 9–14 mm; stigmas not exserted beyond anthers.
subsp. quadrivulnera
Source FNA vol. 10. FNA vol. 10.
Parent taxa Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Biortis Onagraceae > subfam. Onagroideae > tribe Onagreae > Clarkia > sect. Godetia
Sibling taxa
C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. purpurea, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
C. affinis, C. amoena, C. arcuata, C. australis, C. biloba, C. borealis, C. bottae, C. breweri, C. concinna, C. cylindrica, C. davyi, C. delicata, C. dudleyana, C. epilobioides, C. exilis, C. franciscana, C. gracilis, C. heterandra, C. imbricata, C. jolonensis, C. lassenensis, C. lewisii, C. lingulata, C. mildrediae, C. modesta, C. mosquinii, C. prostrata, C. pulchella, C. rhomboidea, C. rostrata, C. rubicunda, C. similis, C. speciosa, C. springvillensis, C. stellata, C. tembloriensis, C. unguiculata, C. virgata, C. williamsonii, C. xantiana
Subordinate taxa
C. purpurea subsp. purpurea, C. purpurea subsp. quadrivulnera, C. purpurea subsp. viminea
Synonyms Oenothera purpurea, Godetia purpurea
Name authority H. Lewis & M. E. Lewis: Madroño 12: 34. (1953) (Curtis) A. Nelson & J. F. Macbride: Bot. Gaz. 65: 64. (1918)
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