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Bolander's claopodium moss

claopodium moss

Habit Plants to 6 cm, in thin mats, green to yellow-green. Plants small to medium-sized, in creeping mats, green to yellowish.
Stem(s)

irregularly branched, smooth;

paraphyllia, when present, squamiform.

and branch leaves similar.;

stem leaves crisped, incurved to slightly contorted when dry, erect and spreading when moist, ovate, ovate-lanceolate, or lanceolate, not plicate;

margins plane, serrate or entire, limbidium sometimes present;

apex gradually or abruptly acuminate, hair-point present or absent;

costa single, ending before apex, pellucid, straight distally;

alar cells not differentiated;

medial and distal laminal cells quadrate-hexagonal, oblong, or rhomboidal, finely and obscurely 1- or multipapillose over lumen, walls moderately thick.

Leaves

ovate, to 1.6 mm;

margins serrate, limbidium indistinctly present;

apex long, narrow, hair-point present;

costa ending before apex, abaxial surface smooth;

distal medial laminal cells short, isodiametric, 6–8 µm, multipapillose.

Seta

1–1.5 cm, rough.

to 2 cm.

Sexual condition

dioicous;

perichaetial leaves pale translucent, longer, apex more acuminate.

Capsule

inclined to horizontal, broadly oval to suborbicular, 1.2–1.8 mm;

operculum long-rostrate.

inclined to horizontal, oblong-oval, asymmetric;

annulus present;

operculum conic- to long-rostrate;

peristome well developed;

exostome teeth lanceolate, densely cross striate-papillose basally, papillose distally;

endostome basal membrane high, segments slender, cilia long.

Spores

12–15 µm, smooth.

9–21 µm, smooth or papillose.

Specialized

asexual reproduction absent.

Claopodium bolanderi

Claopodium

Habitat Rock, soil over rock
Elevation moderate to high elevations
Distribution
from FNA
AK; CA; ID; MT; OR; WA; AB; BC; e Asia (Russia [Commander Islands])
[WildflowerSearch map]
from FNA
HI; w North America; Mexico; w Europe; e Asia
Discussion

Species 8 (4 in the flora).

Claopodium is a genus of small plants with creeping stems, short, papillose laminal cells, pellucid costa, and asymmetric brown capsules with a well-developed peristome. The leaves are broad at the base; the capsules have short necks, are constricted below the mouth when dry, and have bordered exostome teeth and nodose endostome cilia. The genus is somewhat heterogeneous, as C. whippleanum lacks hair-points and superficially is similar to Leskea. Recent phylogenetic work suggests that Claopodium is related to Anomodon (Anomodontaceae).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stem surface cells papillose; laminal cells multipapillose; leaf hair-points present
C. pellucinerve
1. Stem surface cells smooth; laminal cells 1- or multipapillose; leaf hair-points present or absent
→ 2
2. Leaf hair-points absent; laminal cells 1-papillose.
C. whippleanum
2. Leaf hair-points present; laminal cells 1- or multipapillose
→ 3
3. Laminal cells 1-papillose.
C. crispifolium
3. Laminal cells multipapillose.
C. bolanderi
Source FNA vol. 28, p. 344. FNA vol. 28, p. 342. Author: Judith A. Harpel.
Parent taxa Leskeaceae > Claopodium Leskeaceae
Sibling taxa
C. crispifolium, C. pellucinerve, C. whippleanum
Subordinate taxa
C. bolanderi, C. crispifolium, C. pellucinerve, C. whippleanum
Synonyms Hypnum subg. claopodium
Name authority Best: Bull. Torrey Bot. Club 24: 431. (1897) (Lesquereux & James) Renauld & Cardot: Rev. Bryol. 20: 16. (1893)
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