Cistanthe monosperma |
Cistanthe |
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one-seed pussypaws |
pussypaws |
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Habit | Plants perennial, taprooted; caudex unbranched (rarely branching). | Herbs, annual or perennial, succulent, glabrous, with caudices, not rhizomatous or stoloniferous. | ||||||||||||||||||||||||||||||||||||||||
Roots | fleshy and/or fibrous, or taproots. |
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Stems | 2 or more from each rosette, spreading to ascending, to 50 cm. |
decumbent to erect, simple or branched; nodes glabrous. |
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Leaves | basal and cauline; basal in single, prostrate rosette, blade spatulate, 0.5–15 cm; cauline leaves smaller. |
basal and/or cauline, glabrous, not articulate at base, somewhat to markedly clasping, attachment points linear; basal leaves in rosettes; cauline leaves alternate (usually absent in C. umbellata and C. tweedyi). |
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Inflorescences | umbellate, rarely paniculate cymes, usually open. |
axillary, umbellate, paniculate, racemose, or cymose, sometimes scorpioid and/or secund, loose to dense, 1–many-flowered, leafy and/or bracteate; bracts (1–)2 subtending each flower, markedly unequal. |
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Flowers | subsessile; sepals orbiculate to reniform, unequal, 3–8 mm, scarious; petals 4, rose or pinkish white to white, 3–9 mm; stamens 3, anther pink or rose; style exserted, filiform; stigmas 2. |
pedicellate to subsessile; sepals persistent, distinct, not keeled or angled, often unequal, herbaceous to scarious, glabrous; petals 2–9(–12), distinct; stamens 1–23; ovules 1–many; style present or absent; stigmas 2 or 3. |
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Capsules | globose, 2–3.5 mm; valves 2. |
2–3-valved, dehiscence loculicidal from apex, valves not deciduous, (except C. tweedyi, which has circumscissile deshiscence and 3–4 valves splitting from base), margins not reflexed after dehiscence, margins not markedly involute. |
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Seeds | 1–3, black, orbicular-reniform, 0.7–1.2 mm, shiny. |
1–40, dark brownish red, dull gray, or black, usually elliptic to orbiculate, sometimes orbiculate-reniform, dull or shiny, smooth or sculptured, sometimes hairy, strophiolate in C. maritima and C. tweedyi. |
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x | = 22, 23. |
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Cistanthe monosperma |
Cistanthe |
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Phenology | Flowering Apr–Sep. | |||||||||||||||||||||||||||||||||||||||||
Habitat | Dry, sandy or gravelly soils | |||||||||||||||||||||||||||||||||||||||||
Elevation | 300-3500 m (1000-11500 ft) | |||||||||||||||||||||||||||||||||||||||||
Distribution |
CA; NV; OR; Mexico (Baja California)
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South America; North America (including Mexico) |
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Discussion | Species ca. 35 (11 in the flora). Publications by R. C. Carolin (1987) and M. A. Hershkovitz (1991, 1991b) have required the addition to the North American flora of the genus Cistanthe, which originally was erected to accommodate some Chilean species that were segregated from Calandrinia. Recent investigations indicate that a considerably broader range of species belongs in the genus, including two North American species formerly placed in Calandrinia, all the species formerly included in Calyptridium, and one species previously classified in Lewisia. While the current recognition of Cistanthe originally rested on the cladistic work of Carolin and the studies of leaf morphology by Hershkovitz, who documented nine traits of leaf morphology distinguishing Cistanthe, more recent molecular studies by M. A. Hershkovitz and E. A. Zimmer (2000) have provided general support for it. The inclusion of C. tweedyi appears to be somewhat equivocal and it might best be treated as a distinct genus. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 4, p. 463. | FNA vol. 4, p. 460. | ||||||||||||||||||||||||||||||||||||||||
Parent taxa | Portulacaceae > Cistanthe | Portulacaceae | ||||||||||||||||||||||||||||||||||||||||
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Synonyms | Calyptridium monospermum, Spraguea monosperma, Spraguea pulcherrima | Calyptridium, Lewisiopsis, Spraguea | ||||||||||||||||||||||||||||||||||||||||
Name authority | (Greene) Hershkovitz: Phytologia 68: 267. (1990) | Spach: Hist. Nat. Vég. 5: 229. (1836) | ||||||||||||||||||||||||||||||||||||||||
Web links |