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pareira brava, velvetleaf

moonseed family

Stems

striate, without spines.

Leaves

blade 6-12 × 7-14 cm, membranous to nearly leathery;

venation 7.

blade palmately veined, often palmately lobed.

Inflorescences

axillary or terminal, fascicles, cymes, racemes, or panicles, flowers pedicillate.

Flowers

unisexual, staminate and pistillate on different plants, never showy;

sepaloid bracteoles absent;

perianth hypogynous, segments distinct or fused, not showy, greenish white to white or cream, imbricate or valvate.

Staminate flowers

sepals usually 6, not spurred;

petals usually 6, sometimes absent, distinct or connate, ± concave, frequently minute;

nectaries absent;

stamens either opposite petals and equal in number, or numerous;

filaments distinct or united;

anthers dehiscing longitudinally [or transversely];

pistillodes sometimes present.

Pistillate flowers

sepals obovate, 1.2-1.4 × 0.8-1 mm;

petals 4-angulate to deltate, 0.4-0.8 × 1-1.6 mm;

ovary to 0.8 mm.

sepals (4-)6, sometimes reduced to 1;

petals often 6 or reduced to 1, ± concave, usually minute;

nectaries absent;

staminodes frequently present;

pistils 1-6;

ovules 2, aborting to 1, amphitropous;

style often recurved;

stigma entire or lobed.

Fruits

drupes, straight or horseshoe-shaped;

exocarp membranous;

mesocarp ± pulpy;

endocarp (stone) bony, often warty, ribbed.

Drupes

not seen.

Seeds

never stalked;

endosperm present or absent;

embryo usually curved.

Vines

, from thickened root.

and lianas [shrubs or trees], deciduous, woody at least at base, twining or clambering.

Staminate

inflorescences not seen.

Pistillate

inflorescences to 13 cm;

bracts sessile or shortly petiolate, usually reniform, to 5 × 6 mm, grading to minute, membranous, glabrous to densely pilose.

Cissampelos pareira

Menispermaceae

Phenology Flowering summer–winter.
Habitat Hammocks and residential areas
Elevation 0-10 m (0-0 ft)
Distribution
from FNA
FL; Central America; South America; West Indies; Asia; Africa; Pacific Islands
[WildflowerSearch map]
[BONAP county map]
Widespread in temperate and tropical regions
[BONAP county map]
Discussion

Only seven collections of Cissampelos pareira have been seen for North America, all from Dade County, Florida. It is odd that all of them had pistillate flowers. No fruiting material was observed. If staminate plants are present, they are yet to be collected.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genera ca. 78, species ca. 525 (4 genera, 5 species in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Inflorescences with bracts; pistillate flowers: sepal 1, petal 1, pistil 1; staminate flowers (not known in flora): sepals 4, petals 4, connate, stamen filaments fused, anthers 1-locular; leaf blade usually reniform; drupes pubescent
Cissampelos
1. Inflorescences with minute bracts or bracts absent; pistillate and staminate flowers: sepals 6-9, petals distinct when present, stamen filaments distinct, anthers 2-4-locular, pistils 2-6; leaf blade various shapes, rarely reniform; drupes glabrous.
→ 2
2. Petals absent or vestigial; anthers 2-locular; ovary ellipsoid to fusiform; stigma multicleft; leaf blade not mucronate at apex, surfaces sparsely bristly or glabrous; endocarp smooth, excavate on 1 side, margins erose
Calycocarpum
2. Petals well developed; anthers 4-locular; ovary slightly asymmetrically pouched; stigma entire or slightly lobed; leaf blade mucronate at apex, surfaces soft-pubescent or glabrous; endocarp warty and ribbed, depressed but not excavate.
→ 3
3. Inflorescences usually racemes or racemose panicles; petals 6, to 2 mm, with auriculate basal lobes; stamens 6; pistils 6; leaf blades not peltate
Cocculus
3. Inflorescences usually of well-developed panicles; petals (4-)6(-12), to 4 mm, without auriculate basal lobes; stamens 12-36; pistils (2-)3(-4); leaf blade usually peltate, petiole inserted near margin
Menispermum
Source FNA vol. 3. FNA vol. 3, p. 295. Author: Donald G. Rhodes.
Parent taxa Menispermaceae > Cissampelos
Subordinate taxa
Calycocarpum, Cissampelos, Cocculus, Menispermum
Name authority Linnaeus: Sp. Pl. 2: 1031. (1753) Jussieu
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