Flora of North America species comparison

usually 1, erect, glabrous to ± tomentose;

branches many, usually restricted to distal part of stem, ascending.

1, erect, puberulent to pilose with jointed trichomes, sometimes also thinly arachnoid;

branches 0–many, ascending, often nodding at tips.

blades oblong to elliptic, 10–60 × 5–20 cm, unlobed and merely spinulose to irregularly dentate or shallowly to deeply pinnatifid, lobes ± broadly triangular, separated by wide sinuses, obtuse to acute, sometimes coarsely toothed or lobed, main spines slender, 1–3 mm, faces thinly arachnoid, soon glabrescent;

basal often present at flowering, petioles slender, ± winged;

cauline progressively reduced, proximal petiolate, mid and distal sessile, long-decurrent;

distalmost linear to narrowly elliptic, bractlike, spinulose to irregularly dentate or shallowly lobed.

blades oblong to lanceolate or oblanceolate, 10–30 × 2–5 cm, margins flat to undulate, spinulose and otherwise entire to coarsely dentate or shallowly to deeply pinnatifid, lobes well separated, spinulose to coarsely few-dentate, main spines slender to stout, 1–15 mm, one or both faces thinly pilose, sometimes thinly arachnoid, green and ± glabrescent at maturity;

basal usually absent at flowering, sessile or winged-petiolate;

principal cauline well distributed, proximal absent at flowering, moderately reduced distally, winged-petiolate or sessile (proximal), sessile and auriculate-clasping to slightly decurrent 0–2 cm;

distal well developed, spreading, lobed or unlobed.

slender, 1–15 cm.

0–4 cm.

ovoid to hemispheric, 1–2 × 1–2 cm, thinly arachnoid, glabrate.

hemispheric to subspheric, 1.5–2.5 × 1.5–3 cm, glabrous to finely arachnoid and/or pilose, often long pilose-ciliate with arachnoid trichomes connecting adjacent phyllaries.

white to pink-purple, 19–21 mm, tubes 9–10 mm, throats 4–4.5 mm, lobes 5–7 mm;

style tips 2–3.5 mm.

ochroleucous to ± yellow (rarely white or purple), 11–17 mm, tubes 5.5–11 mm, throats 2–4 mm, lobes 3–5 mm;

style tips 2–4 mm.

in 8–9 series, strongly imbricate, green, lanceolate (outer) to linear (inner), abaxial faces with prominent glutinous ridge;

outer and middle appressed, bodies entire, apices acute, spines spreading, slender, ca. 1 mm;

apices of inner often flexuous, acuminate, flat, scabrid-ciliolate.

in 5–8 series, imbricate to subequal, proximally greenish, distally darker, becoming brownish, linear to narrowly lanceolate, outer often nearly as long as inner, abaxial faces with poorly developed glutinous ridge;

outer and mid bases appressed, apices loosely ascending to spreading, bodies entire to spiny-ciliate or terminal appendages expanded, ± scarious, pectinately fringed, spines straight, 2–6 mm;

apices of inner flat or spine-tipped, sometimes expanded and fimbriate.

many, in openly paniculiform arrays, borne singly at tips of peduncles.

1–many, ± erect, loosely to densely clustered at tip of main stem and branches in subcapitate to racemiform arrays, often also in distal leaf axils, closely subtended by clusters of unlobed to deeply dissected, often very spiny bracts.

brown, ca. 4.5 mm, apical collars stramineous, 0.2 mm;

pappi 15–16 mm.

tan to dark brown, 4–6 mm, apical collars narrow, not differently colored;

pappi 9–15 mm.

= 34.