Flora of North America species comparison

1–many, erect or ascending, branches few–many, ascending, villous with septate trichomes.

1–several, usually erect, often stout, glabrous to densely tomentose;

branches 0–many, spreading to ascending, short, stout.

blades oblong-lanceolate to obovate, 15–40 × 6–15 cm, margins plane or revolute, coarsely 1–2-pinnatifid with rigidly divergent lobes, sometimes merely spinose-dentate, lobes triangular to lanceolate, entire to spiny-dentate, main spines 2–10 mm, abaxial faces gray-tomentose, villous with septate trichomes along veins, adaxial green, covered with short appressed bristlelike spines, sometimes tomentose when young;

basal present or absent at flowering, petioles winged, bases tapered;

principal cauline winged-petiolate, mid and distal becoming sessile, well distributed or not, progressively reduced distally, at least distal decurrent as long spiny wings;

distal cauline often more deeply lobed than proximal, main lobes rigidly spiny, margins spinulose, otherwise entire.

blades linear to oblanceolate or oblong-elliptic, 10–40 × 2–10 cm, unlobed and spiny-dentate to deeply pinnatifid, lobes spiny-dentate or coarsely lobed, main spines stout, 5–30 mm, abaxial faces subglabrous to loosely tomentose, adaxial glabrous to ± densely villous with septate trichomes;

basal present at flowering, spiny winged-petiolate, bases often tapered;

principal cauline sessile, well distributed, often not much reduced distally, bases often ± auriculate-clasping;

distal cauline often spinier than the proximal.

1–6 cm.

0–5 cm.

hemispheric to campanulate, 3–4 × 2–4 cm, loosely arachnoid-tomentose.

hemispheric to campanulate, 3–5 × 3–8 cm.

purple (rarely white), 25–35 mm, tubes 18–25 mm, throats 5–6 mm, lobes 5–7 mm;

style tips 3.5–6 mm.

white to yellow, pink, purple, or red, 30–47 mm, tubes 11–30 mm, throats 6–10 mm, lobes 7–10 mm;

style tips 3–5 mm.

in 10–12 series, strongly imbricate, linear-lanceolate (outer) to linear (inner), outer and middle appressed, (bases stramineous), margins entire, abaxial faces without glutinous ridge, apices radiating, greenish, spines 2–5 mm;

apices of inner phyllaries flat, serrulate to minutely erose.

in 5–9 series, subequal to imbricate, light green to stramineous, lanceolate to linear, distally often with reddish margins, abaxial faces without glutinous ridge, often ± thinly tomentose, often scabridulous in submarginal bands;

outer and middle appressed-ascending, bodies usually reddish-tinged, margins setulose-ciliolate, apices acuminate, spines 1–2 mm, weak;

apices of inner straight, flat.

few–many in corymbiform or paniculiform arrays.

1–20 in subcapitate to corymbiform arrays (each closely subtended by an involucre-like ring of spiny-margined bracts).

light brown with darker streaks, 3–4.5 mm, apical collar not differentiated;

pappi 20–30 mm.

straw-colored to tan, 4–6 mm, apical collars weakly differentiated;

pappi 25–35 mm.

= 68.

= 32, 33, 34, 35.