Flora of North America species comparison

1–many, erect or ascending, branches few–many, ascending, villous with septate trichomes.

1–several, erect, thinly gray-tomentose with fine, non-septate trichomes;

branches 0–many, ascending.

blades oblong-lanceolate to obovate, 15–40 × 6–15 cm, margins plane or revolute, coarsely 1–2-pinnatifid with rigidly divergent lobes, sometimes merely spinose-dentate, lobes triangular to lanceolate, entire to spiny-dentate, main spines 2–10 mm, abaxial faces gray-tomentose, villous with septate trichomes along veins, adaxial green, covered with short appressed bristlelike spines, sometimes tomentose when young;

basal present or absent at flowering, petioles winged, bases tapered;

principal cauline winged-petiolate, mid and distal becoming sessile, well distributed or not, progressively reduced distally, at least distal decurrent as long spiny wings;

distal cauline often more deeply lobed than proximal, main lobes rigidly spiny, margins spinulose, otherwise entire.

oblanceolate or elliptic, 15–45 × 2–10 cm, unlobed and merely spinulose to dentate or deeply pinnatifid, lobes well separated, linear to triangular-ovate, merely spinulose to few toothed or lobed near base, margins often revolute, main spines 2–3(–6) mm, abaxial faces densely gray-tomentose, adaxial green, thinly tomentose or ± glabrate;

basal often present at flowering, narrowly winged-petiolate;

principal cauline well distributed, gradually reduced distally, bases of proximal cauline winged-petiolate or sessile, bases of distal cauline expanded and ± clasping, margins sometimes spinier than those of proximal;

distalmost cauline becoming bractlike, often unlobed or less deeply divided than proximal.

1–6 cm.

1–8 cm.

hemispheric to campanulate, 3–4 × 2–4 cm, loosely arachnoid-tomentose.

hemispheric to campanulate, 2.5–3.5 × 2–4 cm, glabrous or loosely floccose.

purple (rarely white), 25–35 mm, tubes 18–25 mm, throats 5–6 mm, lobes 5–7 mm;

style tips 3.5–6 mm.

creamy white, rarely lavender-tinged, 22–28 mm, tubes 8–13 mm, throats 7–11 mm, lobes 4–6 mm;

style tips 5–6 mm.

in 10–12 series, strongly imbricate, linear-lanceolate (outer) to linear (inner), outer and middle appressed, (bases stramineous), margins entire, abaxial faces without glutinous ridge, apices radiating, greenish, spines 2–5 mm;

apices of inner phyllaries flat, serrulate to minutely erose.

in 6–10 series, strongly imbricate, greenish to brown, ovate to lanceolate (outer) to linear (inner), abaxial faces with prominent glutinous ridge;

outer and middle appressed, bodies entire, spines abruptly spreading, fine, 2–3(–5) mm;

apices of inner commonly flexuous or reflexed, flat, scarious.

few–many in corymbiform or paniculiform arrays.

borne singly and terminal on main stems and branches or few from distal axils in corymbiform or paniculiform arrays.

light brown with darker streaks, 3–4.5 mm, apical collar not differentiated;

pappi 20–30 mm.

brown, 5–6 mm, apical collars yellowish, 0.5–1 mm;

pappi 18–22 mm.

= 68.

= 22, 26.