Cirsium turneri |
Cirsium engelmannii |
|
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cliff thistle |
blackland thistle, Engelmann's thistle |
|
Habit | Perennials 15–45 cm; stout, branched caudices. | Biennials or monocarpic perennials, 40–200 cm; taproots and clusters of coarse fibrous roots that often have tuberlike thickenings. |
Stems | 5–30+, horizontal or hanging from cliff sides, thinly appressed gray-tomentose and villous with septate trichomes; branches 0–few, distal, ascending. |
single, erect, often branched above middle, thinly arachnoid-tomentose, ± glabrate; branches few, ascending. |
Leaves | blades oblong-elliptic to oblanceolate, 5–30 × 1–5 cm, shallowly to deeply pinnatifid, lobes spreading, triangular, coarsely dentate or lobed, obtuse to acute, main spines slender, 4–10 mm, abaxial faces green or gray-tomentose, villous with septate trichomes along midveins, ± glabrate, adaxial green and glabrous or thinly tomentose, ± glabrate; basal often present at flowering, spiny winged-petiolate; principal cauline sessile, gradually reduced distally; distal oblong, bases ± clasping, usually less deeply lobed and often spinier than proximal. |
blades elliptic or ovate, 5–20 × 1–10 cm, usually deeply pinnatifid, lobes narrowly to broadly triangular, sinuses broad, rounded (basal and distal cauline sometimes less divided, lobes linear-lanceolate), margins revolute, spreading, entire or spinulose to remotely few-toothed or sharply lobed, main spines slender, 1–5 mm, abaxial faces white-tomentose, adaxial green, villous with septate trichomes or glabrate; basal usually absent at flowering, winged-petiolate, bases tapered; principal cauline well distributed, gradually reduced, bases narrowed, sometimes weakly clasping; distal reduced, widely separated, distalmost bractlike. |
Peduncles | 0–1 cm. |
2–20+ cm, essentially naked with much reduced bracts. |
Involucres | cylindric to narrowly campanulate, 3.5–4.5 × 1.5–2 cm, loosely arachnoid, glabrate, finely short-ciliate. |
ovoid to broadly cylindric or campanulate, 2.5–3.5 × 2–3 cm, thinly arachnoid. |
Corollas | red to reddish purple, 26–27 mm, tubes 3.5–5 mm, throats 7.5–9.5 mm, lobes 12–14 mm; style tips ca. 3 mm. |
pink to purple (white), 32–38 mm, tubes 15–20 mm, throats 6–9 mm, lobes 8–11 mm; style tips 5–6 mm. |
Phyllaries | in 5–6 series, imbricate, linear-lanceolate (outer) to linear (inner), entire, abaxial faces without glutinous ridge, apices red to reddish purple, stiffly ascending, long-acuminate, spines straight, 1–10 mm, ± flattened; apices of inner stramineous to red, straight or flexuous. |
in 10–12 series, strongly imbricate, greenish with subapical darker central zone, ovate (outer) to lanceolate (inner), abaxial faces with narrow glutinous ridge; outer and middle entire, bodies appressed, spines abruptly spreading to deflexed, slender, 2–4 mm; apices of inner phyllaries narrow, flexuous, flattened, entire or finely erose. |
Heads | 1–6+, borne singly or in condensed corymbiform arrays. |
1–10+, borne at tips of main stem and branches. |
Cypselae | stramineous, 5–6 mm, apical collars not differentiated; pappi 20–25 mm. |
brown, 5–6 mm, apical collars yellow, ca. 1 mm; pappi 25–30 mm. |
2n | = 18 (as C. terrae-nigrae), 20 + 1B. |
|
Cirsium turneri |
Cirsium engelmannii |
|
Phenology | Flowering summer (Jun–Sep). | Flowering spring–summer (May–Jul). |
Habitat | Crevices in limestone or basaltic cliffs | Tallgrass prairies, old fields, roadsides, oak savannas, forest edges, in calcareous clay or rarely sandy soils |
Elevation | 900–1500 m (3000–4900 ft) | 50–200 m (200–700 ft) |
Distribution |
TX; Mexico (Chihuahua, Coahuila) |
LA; OK; TX |
Discussion | Cirsium turneri is known from the mountains of the Big Bend area of trans-Pecos Texas and adjacent areas of northern Mexico. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cirsium engelmannii occurs mostly in the blackland prairies of eastern Texas. It ranges northward into southeastern Oklahoma and eastward to northwestern Arkansas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 19, p. 144. | FNA vol. 19, p. 117. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | C. virginianum var. filipendulum, C. terrae-nigrae | |
Name authority | Warnock: SouthW. Naturalist 5: 101. (1960) | Rydberg: Fl. Rocky Mts., 1069. (1917) |
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