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cliff thistle

blackland thistle, Engelmann's thistle

Habit Perennials 15–45 cm; stout, branched caudices. Biennials or monocarpic perennials, 40–200 cm; taproots and clusters of coarse fibrous roots that often have tuberlike thickenings.
Stems

5–30+, horizontal or hanging from cliff sides, thinly appressed gray-tomentose and villous with septate trichomes;

branches 0–few, distal, ascending.

single, erect, often branched above middle, thinly arachnoid-tomentose, ± glabrate;

branches few, ascending.

Leaves

blades oblong-elliptic to oblanceolate, 5–30 × 1–5 cm, shallowly to deeply pinnatifid, lobes spreading, triangular, coarsely dentate or lobed, obtuse to acute, main spines slender, 4–10 mm, abaxial faces green or gray-tomentose, villous with septate trichomes along midveins, ± glabrate, adaxial green and glabrous or thinly tomentose, ± glabrate;

basal often present at flowering, spiny winged-petiolate;

principal cauline sessile, gradually reduced distally;

distal oblong, bases ± clasping, usually less deeply lobed and often spinier than proximal.

blades elliptic or ovate, 5–20 × 1–10 cm, usually deeply pinnatifid, lobes narrowly to broadly triangular, sinuses broad, rounded (basal and distal cauline sometimes less divided, lobes linear-lanceolate), margins revolute, spreading, entire or spinulose to remotely few-toothed or sharply lobed, main spines slender, 1–5 mm, abaxial faces white-tomentose, adaxial green, villous with septate trichomes or glabrate;

basal usually absent at flowering, winged-petiolate, bases tapered;

principal cauline well distributed, gradually reduced, bases narrowed, sometimes weakly clasping;

distal reduced, widely separated, distalmost bractlike.

Peduncles

0–1 cm.

2–20+ cm, essentially naked with much reduced bracts.

Involucres

cylindric to narrowly campanulate, 3.5–4.5 × 1.5–2 cm, loosely arachnoid, glabrate, finely short-ciliate.

ovoid to broadly cylindric or campanulate, 2.5–3.5 × 2–3 cm, thinly arachnoid.

Corollas

red to reddish purple, 26–27 mm, tubes 3.5–5 mm, throats 7.5–9.5 mm, lobes 12–14 mm;

style tips ca. 3 mm.

pink to purple (white), 32–38 mm, tubes 15–20 mm, throats 6–9 mm, lobes 8–11 mm;

style tips 5–6 mm.

Phyllaries

in 5–6 series, imbricate, linear-lanceolate (outer) to linear (inner), entire, abaxial faces without glutinous ridge, apices red to reddish purple, stiffly ascending, long-acuminate, spines straight, 1–10 mm, ± flattened;

apices of inner stramineous to red, straight or flexuous.

in 10–12 series, strongly imbricate, greenish with subapical darker central zone, ovate (outer) to lanceolate (inner), abaxial faces with narrow glutinous ridge;

outer and middle entire, bodies appressed, spines abruptly spreading to deflexed, slender, 2–4 mm;

apices of inner phyllaries narrow, flexuous, flattened, entire or finely erose.

Heads

1–6+, borne singly or in condensed corymbiform arrays.

1–10+, borne at tips of main stem and branches.

Cypselae

stramineous, 5–6 mm, apical collars not differentiated;

pappi 20–25 mm.

brown, 5–6 mm, apical collars yellow, ca. 1 mm;

pappi 25–30 mm.

2n

= 18 (as C. terrae-nigrae), 20 + 1B.

Cirsium turneri

Cirsium engelmannii

Phenology Flowering summer (Jun–Sep). Flowering spring–summer (May–Jul).
Habitat Crevices in limestone or basaltic cliffs Tallgrass prairies, old fields, roadsides, oak savannas, forest edges, in calcareous clay or rarely sandy soils
Elevation 900–1500 m (3000–4900 ft) 50–200 m (200–700 ft)
Distribution
from FNA
TX; Mexico (Chihuahua, Coahuila)
[BONAP county map]
from FNA
LA; OK; TX
[BONAP county map]
Discussion

Cirsium turneri is known from the mountains of the Big Bend area of trans-Pecos Texas and adjacent areas of northern Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cirsium engelmannii occurs mostly in the blackland prairies of eastern Texas. It ranges northward into southeastern Oklahoma and eastward to northwestern Arkansas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 19, p. 144. FNA vol. 19, p. 117.
Parent taxa Asteraceae > tribe Cardueae > Cirsium Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Synonyms C. virginianum var. filipendulum, C. terrae-nigrae
Name authority Warnock: SouthW. Naturalist 5: 101. (1960) Rydberg: Fl. Rocky Mts., 1069. (1917)
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