Cirsium tracyi |
Cirsium mohavense |
|
---|---|---|
Tracy's thistle |
Mohave thistle, Mojave thistle, Rusby's thistle, virgin thistle |
|
Habit | Perennials, 50–200+ cm; tap-rooted. | Biennials or perennials, 30–250 cm; taprooted. |
Stems | 1–several, erect or ascending, thinly gray-tomentose or ± glabrate; branches few to many, ascending. |
1–several, erect, proximally simple, distally branched, ± densely gray-tomentose; branches 0–many, ascending to spreading. |
Leaves | blades elliptic to oblong, 8–40 × 1–12 cm, margins weakly to strongly undulate, spinose-dentate or shallowly to deeply lobed, lobes ascending to spreading, ± triangular, mostly well separated, spinulose and coarsely dentate or cleft into 2–3 lanceolate to triangular, often entire-margined, spine-tipped divisions, main spines 2.5–7+ mm, abaxial faces densely gray-tomentose, adaxial thinly tomentose; basal sometimes present at flowering, winged-petiolate; principal cauline becoming sessile and progressively reduced distally, widest at bases, bases ± auriculate-clasping to short-decurrent; distal cauline reduced, often spinier. |
blades oblong-elliptic to oblanceolate, 10–60 × 2–15 cm, unlobed and merely spinulose or spiny-dentate or shallowly to deeply pinnatifid, lobes linear-lanceolate to ovate-triangular, spreading, entire to coarsely dentate, main spines slender to stout, 3–30 mm, faces ± gray-tomentose, sometimes ± glabrate; basal often present at flowering, winged-petiolate; principal cauline decreasing distally, proximal winged-petiolate, distal sessile, bases decurrent as spiny wings 1–5 cm; distalmost well separated, bractlike. |
Peduncles | 0–10+ cm. |
0–15 cm. |
Involucres | ovoid to hemispheric or broadly campanulate, 2–3 × 1.7–3.5 cm, loosely arachnoid on phyllary margins or glabrate. |
ovoid to hemispheric, 1.5–2.5 × 1.5–2 cm, loosely arachnoid on phyllary margins or glabrate. |
Corollas | white to lavender or pink-purple, 23–30 mm, tubes 9–14 mm, throats 5.5–10.5 mm, lobes 5.5–9.5 mm; style tips 4–7 mm. |
white to pink or lavender, 16–25 mm, tubes 7–12 mm, throats 4–7 mm, lobes 4–8 mm, style tips 3–4 mm. |
Phyllaries | in 6–10 series, imbricate, ovate to lanceolate (outer) to linear-lanceolate (inner), margins entire, abaxial faces with prominent glutinous ridge; outer and middle appressed, spines spreading, slender to stout, 2–6 mm; apices of inner often flexuous, narrow, flat, ± entire, spineless or tipped with weak spines. |
in 5–8 series, imbricate, (inner greenish to brown or stramineous), lanceolate or ovate (outer) to linear-lanceolate (inner), entire, abaxial faces with narrow glutinous ridge; outer and middle appressed, spines spreading, 3–7 mm; apices of inner often flexuous, flattened, spineless, scabrid. |
Heads | 1–many, terminal on branches and often in leaf axils, in leafy, ± corymbiform arrays. |
1–many, in corymbiform or paniculiform arrays. |
Cypselae | light to dark brown, 6–7 mm, apical collars colored like body or rarely yellowish, narrow; pappi 20–23 mm, usually noticeably shorter than corolla. |
stramineous to dark brown, 3–6 mm, apical collars 0.2–0.3 mm, yellowish; pappi 14–16 mm. |
2n | = 24. |
= 30, 32. |
Cirsium tracyi |
Cirsium mohavense |
|
Phenology | Flowering late spring–summer (Jun–Aug). | Flowering summer–fall (Jun–Oct). |
Habitat | Dry slopes, sagebrush deserts, pinyon-juniper woodlands, openings in montane coniferous forests, often in disturbed areas | Wet soil, streams, springs, meadows in desert and desert woodland areas |
Elevation | 1400–2900 m (4600–9500 ft) | -50–2200 m (-200–7200 ft) |
Distribution |
CO; NM; UT
|
AZ; CA; NV; UT
|
Discussion | Cirsium tracyi occurs from eastern Utah and western Colorado south in the Colorado Plateau and southern Rocky Mountains to northwestern New Mexico. Large-headed plants of Cirsium tracyi and small-headed individuals of C. undulatum are sometimes difficult to distinguish. P. L. Barlow-Irick (unpubl.) found that although there is much overlap in floral measurements of C. tracyi and C. undulatum, the means for some of these characters are statistically significant. Corolla lobes of C. tracyi, for instance, average about 7 mm and those of C. undulatum about 10 mm. The species differ in chromosome number as well. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Cirsium mohavense ranges from scattered sites in eastern California east in the Basin and Range Province of southern Nevada to southwestern Utah and nortwestern Arizona, mostly in Mojave Desert region. When Welsh proposed Cirsium virginense for a geographically limited group of plants from southwestern Utah and northwestern Arizona (and subsequently discovered in extreme southeastern Nevada), he indicated that its relationship to other western thistles was unknown. Subsequently, he indicated (S. L. Welsh 1983; Welsh et al. 1993) that the affinities of the taxon apparently lie with C. mohavense, but he did not attempt to distinguish C. virginense from C. mohavense (in the strict sense) because the latter was not known to occur in Utah. A. Cronquist (1994) attempted the distinction. The only character he used in his key was life span of the plants: biennial (C. mohavense) versus perennial, spreading by creeping roots (C. virginense). In the descriptions of the two taxa he elaborated on this character, indicating that C. mohavense is single-stemmed and C. virginense often multistemmed. In the remaining features the plants are very similar or overlap extensively. Distinction of two taxa on the basis of duration is impractical and probably inaccurate. Specimens commonly lack roots, and in those specimens in which bases are present, I have seldom been able to make any distinction between biennial taproots and perennial taproots. In particular I have seen no evidence of creeping roots. I am not aware of any study of either taxon that documents the life history of the plants. Some specimens of C. mohavense (in the strict sense) appear to have perennial bases like those attributed to C. virginense by Cronquist. For instance, a specimen of C. mohavense from Death Valley (Thorne & Ratcliff 2287, BRY) is indistinguishable from specimens of C. virginense (e.g., Atwood 13374, BRY) from Nevada and Utah. Both have a branched root crown with multiple rosettes and nearly identical leaves and heads. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 19, p. 121. | FNA vol. 19, p. 134. |
Parent taxa | Asteraceae > tribe Cardueae > Cirsium | Asteraceae > tribe Cardueae > Cirsium |
Sibling taxa | ||
Synonyms | Carduus tracyi, C. acuatum, C. floccosum, C. undulatus var. tracyi | Carduus mohavensis, C. rusbyi, C. virginense |
Name authority | (Rydberg) Petrak: Beih. Bot. Centralbl. 35(2): 424. (1917) | (Greene) Petrak: Bot. Tidsskr. 31: 68. (1911) |
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