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Tracy's thistle

blackland thistle, Engelmann's thistle

Habit Perennials, 50–200+ cm; tap-rooted. Biennials or monocarpic perennials, 40–200 cm; taproots and clusters of coarse fibrous roots that often have tuberlike thickenings.
Stems

1–several, erect or ascending, thinly gray-tomentose or ± glabrate;

branches few to many, ascending.

single, erect, often branched above middle, thinly arachnoid-tomentose, ± glabrate;

branches few, ascending.

Leaves

blades elliptic to oblong, 8–40 × 1–12 cm, margins weakly to strongly undulate, spinose-dentate or shallowly to deeply lobed, lobes ascending to spreading, ± triangular, mostly well separated, spinulose and coarsely dentate or cleft into 2–3 lanceolate to triangular, often entire-margined, spine-tipped divisions, main spines 2.5–7+ mm, abaxial faces densely gray-tomentose, adaxial thinly tomentose;

basal sometimes present at flowering, winged-petiolate;

principal cauline becoming sessile and progressively reduced distally, widest at bases, bases ± auriculate-clasping to short-decurrent;

distal cauline reduced, often spinier.

blades elliptic or ovate, 5–20 × 1–10 cm, usually deeply pinnatifid, lobes narrowly to broadly triangular, sinuses broad, rounded (basal and distal cauline sometimes less divided, lobes linear-lanceolate), margins revolute, spreading, entire or spinulose to remotely few-toothed or sharply lobed, main spines slender, 1–5 mm, abaxial faces white-tomentose, adaxial green, villous with septate trichomes or glabrate;

basal usually absent at flowering, winged-petiolate, bases tapered;

principal cauline well distributed, gradually reduced, bases narrowed, sometimes weakly clasping;

distal reduced, widely separated, distalmost bractlike.

Peduncles

0–10+ cm.

2–20+ cm, essentially naked with much reduced bracts.

Involucres

ovoid to hemispheric or broadly campanulate, 2–3 × 1.7–3.5 cm, loosely arachnoid on phyllary margins or glabrate.

ovoid to broadly cylindric or campanulate, 2.5–3.5 × 2–3 cm, thinly arachnoid.

Corollas

white to lavender or pink-purple, 23–30 mm, tubes 9–14 mm, throats 5.5–10.5 mm, lobes 5.5–9.5 mm;

style tips 4–7 mm.

pink to purple (white), 32–38 mm, tubes 15–20 mm, throats 6–9 mm, lobes 8–11 mm;

style tips 5–6 mm.

Phyllaries

in 6–10 series, imbricate, ovate to lanceolate (outer) to linear-lanceolate (inner), margins entire, abaxial faces with prominent glutinous ridge;

outer and middle appressed, spines spreading, slender to stout, 2–6 mm;

apices of inner often flexuous, narrow, flat, ± entire, spineless or tipped with weak spines.

in 10–12 series, strongly imbricate, greenish with subapical darker central zone, ovate (outer) to lanceolate (inner), abaxial faces with narrow glutinous ridge;

outer and middle entire, bodies appressed, spines abruptly spreading to deflexed, slender, 2–4 mm;

apices of inner phyllaries narrow, flexuous, flattened, entire or finely erose.

Heads

1–many, terminal on branches and often in leaf axils, in leafy, ± corymbiform arrays.

1–10+, borne at tips of main stem and branches.

Cypselae

light to dark brown, 6–7 mm, apical collars colored like body or rarely yellowish, narrow;

pappi 20–23 mm, usually noticeably shorter than corolla.

brown, 5–6 mm, apical collars yellow, ca. 1 mm;

pappi 25–30 mm.

2n

= 24.

= 18 (as C. terrae-nigrae), 20 + 1B.

Cirsium tracyi

Cirsium engelmannii

Phenology Flowering late spring–summer (Jun–Aug). Flowering spring–summer (May–Jul).
Habitat Dry slopes, sagebrush deserts, pinyon-juniper woodlands, openings in montane coniferous forests, often in disturbed areas Tallgrass prairies, old fields, roadsides, oak savannas, forest edges, in calcareous clay or rarely sandy soils
Elevation 1400–2900 m (4600–9500 ft) 50–200 m (200–700 ft)
Distribution
from FNA
CO; NM; UT
[WildflowerSearch map]
[BONAP county map]
from FNA
LA; OK; TX
[BONAP county map]
Discussion

Cirsium tracyi occurs from eastern Utah and western Colorado south in the Colorado Plateau and southern Rocky Mountains to northwestern New Mexico. Large-headed plants of Cirsium tracyi and small-headed individuals of C. undulatum are sometimes difficult to distinguish. P. L. Barlow-Irick (unpubl.) found that although there is much overlap in floral measurements of C. tracyi and C. undulatum, the means for some of these characters are statistically significant. Corolla lobes of C. tracyi, for instance, average about 7 mm and those of C. undulatum about 10 mm. The species differ in chromosome number as well.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Cirsium engelmannii occurs mostly in the blackland prairies of eastern Texas. It ranges northward into southeastern Oklahoma and eastward to northwestern Arkansas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 19, p. 121. FNA vol. 19, p. 117.
Parent taxa Asteraceae > tribe Cardueae > Cirsium Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Synonyms Carduus tracyi, C. acuatum, C. floccosum, C. undulatus var. tracyi C. virginianum var. filipendulum, C. terrae-nigrae
Name authority (Rydberg) Petrak: Beih. Bot. Centralbl. 35(2): 424. (1917) Rydberg: Fl. Rocky Mts., 1069. (1917)
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