Cirsium tracyi |
Cirsium crassicaule |
|
---|---|---|
Tracy's thistle |
slough thistle |
|
Habit | Perennials, 50–200+ cm; tap-rooted. | Annuals or biennials, (60–)100–300 cm; taprooted. |
Stems | 1–several, erect or ascending, thinly gray-tomentose or ± glabrate; branches few to many, ascending. |
usually 1, erect, stout, (hollow, 2–10 cm diam. at base), openly branched distally, thinly arachnoid, villous with jointed trichomes, at least proximally. |
Leaves | blades elliptic to oblong, 8–40 × 1–12 cm, margins weakly to strongly undulate, spinose-dentate or shallowly to deeply lobed, lobes ascending to spreading, ± triangular, mostly well separated, spinulose and coarsely dentate or cleft into 2–3 lanceolate to triangular, often entire-margined, spine-tipped divisions, main spines 2.5–7+ mm, abaxial faces densely gray-tomentose, adaxial thinly tomentose; basal sometimes present at flowering, winged-petiolate; principal cauline becoming sessile and progressively reduced distally, widest at bases, bases ± auriculate-clasping to short-decurrent; distal cauline reduced, often spinier. |
blades elliptic to broadly oblanceolate, 15–70 × 30–150+ cm, flat, pinnatifid 1/2–2/3 distance to midvein, larger usually with broad sinuses, lobes broad, few lobed or dentate, main spines 3–8 mm, abaxial faces gray-tomentose, adaxial thinly arachnoid-tomentose, sometimes midveins with jointed trichomes; basal present or withered at flowering, winged-petiolate; principal cauline sessile, progressively reduced distally, bases clasping or short-decurrent 1–2 cm; distal cauline reduced, becoming bractlike, sometimes spinier than proximal. |
Peduncles | 0–10+ cm. |
0–15 cm. |
Involucres | ovoid to hemispheric or broadly campanulate, 2–3 × 1.7–3.5 cm, loosely arachnoid on phyllary margins or glabrate. |
ovoid to campanulate, 1.5–3 × 1.5–3 cm, ± glabrous. |
Corollas | white to lavender or pink-purple, 23–30 mm, tubes 9–14 mm, throats 5.5–10.5 mm, lobes 5.5–9.5 mm; style tips 4–7 mm. |
pale rose-purple (white), 19–26 mm, tubes 9–12 mm, throats 4–6 mm, lobes 5–9 mm; style tips 3.5–4.5 mm. |
Phyllaries | in 6–10 series, imbricate, ovate to lanceolate (outer) to linear-lanceolate (inner), margins entire, abaxial faces with prominent glutinous ridge; outer and middle appressed, spines spreading, slender to stout, 2–6 mm; apices of inner often flexuous, narrow, flat, ± entire, spineless or tipped with weak spines. |
in 5–7 series, weakly unequal, dark green to brownish, lanceolate (outer) to linear (inner), abaxial faces without (or with very obscure) glutinous ridge; outer and middle appressed or apices spreading, at least outer irregularly spiny-fringed, finely serrulate, spines slender, 3–5 mm; apices of inner erect, abaxial faces gray-tomentose, ± twisted. |
Heads | 1–many, terminal on branches and often in leaf axils, in leafy, ± corymbiform arrays. |
1–several at branch tips, closely subtended by clustered leafy bracts or not, collectively forming open, corymbiform or paniculiform arrays. |
Cypselae | light to dark brown, 6–7 mm, apical collars colored like body or rarely yellowish, narrow; pappi 20–23 mm, usually noticeably shorter than corolla. |
dark brown, 5–5.5 mm, collars narrow, ± stramineous; pappi 15–20 mm. |
2n | = 24. |
= 32. |
Cirsium tracyi |
Cirsium crassicaule |
|
Phenology | Flowering late spring–summer (Jun–Aug). | Flowering spring (Apr–Jun). |
Habitat | Dry slopes, sagebrush deserts, pinyon-juniper woodlands, openings in montane coniferous forests, often in disturbed areas | Freshwater marshes, canal banks |
Elevation | 1400–2900 m (4600–9500 ft) | 5–100 m (0–300 ft) |
Distribution |
CO; NM; UT
|
CA |
Discussion | Cirsium tracyi occurs from eastern Utah and western Colorado south in the Colorado Plateau and southern Rocky Mountains to northwestern New Mexico. Large-headed plants of Cirsium tracyi and small-headed individuals of C. undulatum are sometimes difficult to distinguish. P. L. Barlow-Irick (unpubl.) found that although there is much overlap in floral measurements of C. tracyi and C. undulatum, the means for some of these characters are statistically significant. Corolla lobes of C. tracyi, for instance, average about 7 mm and those of C. undulatum about 10 mm. The species differ in chromosome number as well. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cirsium crassicaule is known only from a few sites in the San Joaquin Valley. Some populations are threatened by habitat modification and development pressures. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 19, p. 121. | FNA vol. 19, p. 132. |
Parent taxa | Asteraceae > tribe Cardueae > Cirsium | Asteraceae > tribe Cardueae > Cirsium |
Sibling taxa | ||
Synonyms | Carduus tracyi, C. acuatum, C. floccosum, C. undulatus var. tracyi | Carduus crassicaulis |
Name authority | (Rydberg) Petrak: Beih. Bot. Centralbl. 35(2): 424. (1917) | (Greene) Jepson: Fl. W. Calif., 506. (1901) |
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