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chardon écailleux, dinnerplate thistle, elk thistle, meadow thistle

Beaumont thistle, yellowspine thistle

Habit Biennials or monocarpic perennials, acaulescent, short caulescent and forming low rounded mounds, or caulescent and erect, 0–200 cm; taprooted. Perennials, 30–90 cm; crown sprouts or runner roots producing adventitious buds.
Stems

absent, or with crowded branches from near base, or simple and erect, often fleshy and thickened, glabrous to thinly gray- tomentose, often villous with septate trichomes.

1–20+, erect or ascending, densely gray-tomentose with non-septate trichomes;

branches 0 or few, usually in distal 1/2, ascending.

Leaves

blades linear to elliptic, 5–20 × 3–7 cm, plane to strongly undulate, unlobed or shallowly to deeply pinnatifid, lobes linear-lanceolate to broadly triangular, closely spaced, spreading, spinose-dentate or lobed, main spines slender to stout, 2–15+ mm, abaxial faces glabrous or thinly to densely tomentose, ± villous with septate trichomes along the veins, glabrate or trichomes persistent, adaxial thinly arachnoid tomentose and soon glabrescent;

basal often present at flowering, sessile or winged-petiolate;

cauline many in caulescent forms, reduced distally or not, winged-petiolate or distal sessile;

distal often well developed, similar to proximal, sometimes much narrower and bractlike.

blades oblong to narrowly elliptic, 10–30 × 2–8 cm, strongly undulate, margins coarsely dentate or shallowly to deeply pinnatifid with 8–15 pairs of lobes 0.5–2 cm, often revolute, lobes ± triangular, closely spaced, spreading, spinose-dentate and cleft into 2–5 spine-tipped divisions, main spines 5–20 mm, yellowish, abaxial faces densely white-tomentose, adaxial thinly gray-tomentose;

basal usually present at flowering, winged-petiolate;

principal cauline sessile, progressively reduced distally, bases ± auriculate to long-decurrent as spiny wings;

distal cauline usually much reduced, less lobed.

Peduncles

0–10 cm, leafy-bracted.

0–4 cm.

Involucres

ovoid to hemispheric, 2–4 × 1.5–6 cm, loosely arachnoid on phyllary margins or glabrate.

ovoid to hemispheric or broadly campanulate, 2.5–4.5 × 2.5–4.5 cm in first-formed heads, often smaller in later ones, loosely arachnoid on phyllary margins or glabrate.

Corollas

white or pale lavender to purple, 20–40 mm, tubes 7–24 mm, throats 4–12 mm (noticeably larger than tubes), lobes 4–10 mm;

style tips 3.5–8 mm.

white or pale lavender to purple, pink, or red, 25–45 mm, tubes 8–25 mm, throats 6–17 mm, lobes 6–15 mm;

style tips 2–8 mm.

Phyllaries

in 5–10 series, imbricate, ovate or lanceolate (outer) to linear or linear-lanceolate (inner), margins (outer) entire or scarious-fringed, abaxial faces without glutinous ridge;

outer and mid appressed, spines erect to spreading 0.5–13 mm;

apices of mid and inner narrowed and scabro-denticulate or with expanded, erose-dentate tips, spineless or tipped with flattened spines.

in 5–10 series, imbricate, ovate (outer) to linear-lanceolate (inner), margins entire, abaxial faces with narrow glutinous ridge;

outer and middle appressed, spines spreading, 3–12 mm;

apices of inner often flexuous, expanded and flat, scabrid-margined, sometimes erose, spineless.

Heads

1–many, erect, borne singly or often densely crowded in spiciform, racemiform, or subcapitate arrays, especially in acaulescent or short-caulescent plants, often closely subtended by distalmost leaves.

1–few, in leafy, ± corymbiform arrays.

Cypselae

light to dark brown, 4–6.5 mm, apical collars usually colored like body;

pappi 17–35 mm, white to tan.

light brown, sometimes with lighter or darker streaks, 6–9 mm, apical collars colored like the body, narrow;

pappi (white or tawny), 20–40 mm, usually noticeably shorter than corolla.

2n

= 34, 36.

= 30, 31, 32, 34.

Cirsium scariosum

Cirsium ochrocentrum

Distribution
from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; QC; disjunct to e Que (Mingan Archipelago)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; KS; NE; NM; OK; SD; TX; UT; WY; n Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 8 (8 in the flora).

Cirsium scariosum is a widely distributed complex of intergrading races distributed from southwestern Canada to northwestern Mexico. These plants range from acaulescent rosettes with a tight cluster of sessile heads to tall, erect, unbranched plants, or moundlike, more or less openly branched herbs. Acaulescent and caulescent plants sometimes occur in the same population.

Members of this complex have been variously treated in the past. F. Petrak (1917) recognized ten species plus several subspecies for the taxa I am treating here as C. scariosum (in the broad sense). In floras, the names C. drummondii and C. foliosum have been widely misapplied to these plants (R. J. Moore and C. Frankton 1964). The latter two species, while clearly related to C. scariosum, have a range restricted mostly to Canada. Moore and Frankton (1967) attempted to bring order to the complex and recognized four species for plants that I include here in C. scariosum: C. acaulescens, C. congdonii, C. coloradense, and C. scariosum in the restricted sense. Moore and Frankton substituted the prior name C. tioganum for C. acaulescens. Unfortunately they did not extend their study widely enough and did not include some members of the complex in their investigations. S. L. Welsh (1982) proposed C. scariosum var. thorneae from Utah and lumped the various species recognized by Moore and Frankton within a highly polymorphic var. scariosum. After consulting with A. Cronquist and studying his manuscript treatment of Cirsium for the Intermountain Flora, D. J. Keil and C. E. Turner (1993) also accepted a broadly construed C. scariosum. Cronquist (1994) treated C. scariosum as an extremely variable species that included the four species recognized by Moore and Frankton plus the variety proposed by Welsh. Cronquist chose to not recognize infraspecific taxa.

In the present treatment I have examined these plants from a biogeographic perspective with the goal of discerning regional patterns of variation. The large number of specimens available has allowed me to examine distributional patterns in relation to the topography and biogeographic history of the regions where this species occurs. My field studies also have provided me with observations that help to explain some of the anomalous specimens represented in herbaria. Although the variation within and between populations is sometimes amazing, more-or-less differentiated geographic races can be discerned. Because of the extraordinary and overlapping patterns of variation across the range of Cirsium scariosum, the following key to varieties should be regarded as at best an approximation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants acaulescent (occasional short-caulescent individuals sometimes present in a population)
→ 2
1. Plants caulescent (occasional acaulescent individuals sometimes present in a population)
→ 4
2. Corollas pink to purple; Sierra Nevada of w Nevada and e California to San Bernardino Mountains of s California
var. congdonii
2. Corollas white to faintly pink-or lilac-tinged; widespread
→ 3
3. Abaxial leaf faces usually gray-tomentose; widespread, Colorado to s Oregon, n California
var. americanum
3. Abaxial leaf faces usually green, glabrous or glabrate; s. California
var. citrinum
4. Larger leaf spines 1–3 cm
Cirsium scariosum var. thorneae
4. Larger leaf spines usually shorter
→ 5
5. Corollas purple
→ 6
5. Corollas white to faintly pink- or lilac-tinged
→ 7
6. Corolla lobes 5.5–8 mm; sw. Idaho, n Nevada, se Oregon
var. toiyabense
6. Corolla lobes 3.5–6 mm; e Oregon to sw Montana
var. scariosum
7. Stems usually proximally branched, plants often forming low, rounded mound; heads usually borne on short to ± elongate lateral branches; corollas 26–36 mm
→ 8
7. Stems usually erect, proximally unbranched; heads usually sessile or short-pedunculate in subcapitate to spiciform or racemiform arrays, usually closely subtended by numerous distal leaves; corollas 20–29 mm
→ 9
8. Apices of inner phyllaries acuminate and entire or rarely toothed; s California
var. citrinum
8. Apices of inner phyllaries usually expanded as a scarious, erose-toothed appendage; ne California, se Oregon
var. robustum
9. Heads usually ± tightly clustered at stem tips, closely subtended and often overtopped by crowded distal leaves; distal leaves ± thin, usually fringed with numerous weak spines, often ± unpigmented proximally or tinged pink to purplish
var. scariosum
9. Heads usually in ± leafy, racemiform arrays, usually subtended by ± reduced, bractlike distal leaves; distal leaves firm, strongly spiny, usually green throughout
var. coloradense
1. Stems densely leafy, nodes crowded; leaves often long- decurrent; corollas white or pale lavender to purple
var. ochrocentrum
1. Stems leafy, nodes usually well separated; distal cauline leaves clasping, or if decurrent spiny wing usually less than 1 cm; corollas red, pink, or reddish purple (rarely white)
var. martinii
Source FNA vol. 19, p. 1. FNA vol. 19, p. 123.
Parent taxa Asteraceae > tribe Cardueae > Cirsium Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Subordinate taxa
C. scariosum var. americanum, C. scariosum var. citrinum, C. scariosum var. coloradense, C. scariosum var. congdonii, C. scariosum var. robustum, C. scariosum var. scariosum, C. scariosum var. toiyabense
C. ochrocentrum var. martinii, C. ochrocentrum var. ochrocentrum
Synonyms C. hookerianum var. scariosum
Name authority Nuttall: Trans. Amer. Philos. Soc., n. s. 7: 420. (1841) A. Gray: Mem. Amer. Acad. Arts, n. s. 4: 110. (1849)
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