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chardon écailleux, dinnerplate thistle, elk thistle, meadow thistle

graygreen thistle, peregrine thistle

Habit Biennials or monocarpic perennials, acaulescent, short caulescent and forming low rounded mounds, or caulescent and erect, 0–200 cm; taprooted. Biennials or perennials, 25–120 cm, pubescence a mixture of fine, non-septate arachnoid trichomes and coarser, septate trichomes, especially along stems and on midveins on abaxial leaf faces, usually ± loose and irregularly deciduous from leaves in age; taprooted.
Stems

absent, or with crowded branches from near base, or simple and erect, often fleshy and thickened, glabrous to thinly gray- tomentose, often villous with septate trichomes.

usually 1, erect, ± gray-tomentose, sometimes villous with septate trichomes;

branches 0–10+, usually arising in distal 1/2, ascending, usually reaching a ± common height.

Leaves

blades linear to elliptic, 5–20 × 3–7 cm, plane to strongly undulate, unlobed or shallowly to deeply pinnatifid, lobes linear-lanceolate to broadly triangular, closely spaced, spreading, spinose-dentate or lobed, main spines slender to stout, 2–15+ mm, abaxial faces glabrous or thinly to densely tomentose, ± villous with septate trichomes along the veins, glabrate or trichomes persistent, adaxial thinly arachnoid tomentose and soon glabrescent;

basal often present at flowering, sessile or winged-petiolate;

cauline many in caulescent forms, reduced distally or not, winged-petiolate or distal sessile;

distal often well developed, similar to proximal, sometimes much narrower and bractlike.

blades linear-oblong to oblanceolate or elliptic, 10–30 × 3–7 cm, shallowly to deeply pinnatifid with 3–8 pairs of lobes, longer than 2 cm, lobes well separated, linear to triangular-ovate, dentate to lobed proximally, main spines slender, 2–7 mm, faces green to gray, thinly to densely arachnoid-tomentose with fine, non-septate trichomes, sometimes villous with septate trichomes along veins, usually ± loose and irregularly deciduous from leaves in age;

basal often present at flowering, sessile or winged-petiolate;

principal cauline mostly in proximal 1/2, winged-petiolate or sessile, bases narrowed, auriculate, veins often prominently raised on abaxial faces;

distal sessile, auriculate-clasping or short-decurrent 1–10 mm, progressively reduced becoming bractlike, often unlobed or less deeply divided and sometimes spinier than proximal.

Peduncles

0–10 cm, leafy-bracted.

(0–)2–15 cm.

Involucres

ovoid to hemispheric, 2–4 × 1.5–6 cm, loosely arachnoid on phyllary margins or glabrate.

ovoid to hemispheric or campanulate, 2–3 × 1.5–3.5 cm, ± arachnoid-floccose, often glabrate.

Corollas

white or pale lavender to purple, 20–40 mm, tubes 7–24 mm, throats 4–12 mm (noticeably larger than tubes), lobes 4–10 mm;

style tips 3.5–8 mm.

creamy white to purplish, 20–31 mm, tubes 8–14 mm, throats 5.5–10 mm, lobes 6–7 mm;

style tips 4–6 mm.

Phyllaries

in 5–10 series, imbricate, ovate or lanceolate (outer) to linear or linear-lanceolate (inner), margins (outer) entire or scarious-fringed, abaxial faces without glutinous ridge;

outer and mid appressed, spines erect to spreading 0.5–13 mm;

apices of mid and inner narrowed and scabro-denticulate or with expanded, erose-dentate tips, spineless or tipped with flattened spines.

in 8–10 series, subequal to strongly imbricate, green, linear to lanceolate (outer) to linear (inner), entire, abaxial faces with inconspicuous to prominent glutinous ridge;

outer and mid bodies loosely spreading to ascending or appressed, apices subappressed to ascending or spreading, flat, spines ascending to spreading, fine, 2–4 mm;

apices of inner commonly flexuous or reflexed, narrow, flat, scarious.

Heads

1–many, erect, borne singly or often densely crowded in spiciform, racemiform, or subcapitate arrays, especially in acaulescent or short-caulescent plants, often closely subtended by distalmost leaves.

borne singly, terminal on main stem and branches, sometimes also in distal axils, erect, not subtended by well-developed leaves, collectively forming corymbiform or racemiform arrays.

Cypselae

light to dark brown, 4–6.5 mm, apical collars usually colored like body;

pappi 17–35 mm, white to tan.

tan to dark brown, 5–7.5 mm, apical collars not differentiated;

pappi 16–25 mm.

2n

= 34, 36.

Cirsium scariosum

Cirsium cymosum

Distribution
from FNA
AZ; CA; CO; ID; MT; NM; NV; OR; UT; WA; WY; AB; BC; QC; disjunct to e Que (Mingan Archipelago)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; ID; MT; NV; OR; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 8 (8 in the flora).

Cirsium scariosum is a widely distributed complex of intergrading races distributed from southwestern Canada to northwestern Mexico. These plants range from acaulescent rosettes with a tight cluster of sessile heads to tall, erect, unbranched plants, or moundlike, more or less openly branched herbs. Acaulescent and caulescent plants sometimes occur in the same population.

Members of this complex have been variously treated in the past. F. Petrak (1917) recognized ten species plus several subspecies for the taxa I am treating here as C. scariosum (in the broad sense). In floras, the names C. drummondii and C. foliosum have been widely misapplied to these plants (R. J. Moore and C. Frankton 1964). The latter two species, while clearly related to C. scariosum, have a range restricted mostly to Canada. Moore and Frankton (1967) attempted to bring order to the complex and recognized four species for plants that I include here in C. scariosum: C. acaulescens, C. congdonii, C. coloradense, and C. scariosum in the restricted sense. Moore and Frankton substituted the prior name C. tioganum for C. acaulescens. Unfortunately they did not extend their study widely enough and did not include some members of the complex in their investigations. S. L. Welsh (1982) proposed C. scariosum var. thorneae from Utah and lumped the various species recognized by Moore and Frankton within a highly polymorphic var. scariosum. After consulting with A. Cronquist and studying his manuscript treatment of Cirsium for the Intermountain Flora, D. J. Keil and C. E. Turner (1993) also accepted a broadly construed C. scariosum. Cronquist (1994) treated C. scariosum as an extremely variable species that included the four species recognized by Moore and Frankton plus the variety proposed by Welsh. Cronquist chose to not recognize infraspecific taxa.

In the present treatment I have examined these plants from a biogeographic perspective with the goal of discerning regional patterns of variation. The large number of specimens available has allowed me to examine distributional patterns in relation to the topography and biogeographic history of the regions where this species occurs. My field studies also have provided me with observations that help to explain some of the anomalous specimens represented in herbaria. Although the variation within and between populations is sometimes amazing, more-or-less differentiated geographic races can be discerned. Because of the extraordinary and overlapping patterns of variation across the range of Cirsium scariosum, the following key to varieties should be regarded as at best an approximation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Past floras have treated Cirsium cymosum and C. canovirens as separate species. In my examination of these plants across their combined ranges I realized that they are connected by numerous intermediates and that I could find no characters that consistently distinguish them.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants acaulescent (occasional short-caulescent individuals sometimes present in a population)
→ 2
1. Plants caulescent (occasional acaulescent individuals sometimes present in a population)
→ 4
2. Corollas pink to purple; Sierra Nevada of w Nevada and e California to San Bernardino Mountains of s California
var. congdonii
2. Corollas white to faintly pink-or lilac-tinged; widespread
→ 3
3. Abaxial leaf faces usually gray-tomentose; widespread, Colorado to s Oregon, n California
var. americanum
3. Abaxial leaf faces usually green, glabrous or glabrate; s. California
var. citrinum
4. Larger leaf spines 1–3 cm
Cirsium scariosum var. thorneae
4. Larger leaf spines usually shorter
→ 5
5. Corollas purple
→ 6
5. Corollas white to faintly pink- or lilac-tinged
→ 7
6. Corolla lobes 5.5–8 mm; sw. Idaho, n Nevada, se Oregon
var. toiyabense
6. Corolla lobes 3.5–6 mm; e Oregon to sw Montana
var. scariosum
7. Stems usually proximally branched, plants often forming low, rounded mound; heads usually borne on short to ± elongate lateral branches; corollas 26–36 mm
→ 8
7. Stems usually erect, proximally unbranched; heads usually sessile or short-pedunculate in subcapitate to spiciform or racemiform arrays, usually closely subtended by numerous distal leaves; corollas 20–29 mm
→ 9
8. Apices of inner phyllaries acuminate and entire or rarely toothed; s California
var. citrinum
8. Apices of inner phyllaries usually expanded as a scarious, erose-toothed appendage; ne California, se Oregon
var. robustum
9. Heads usually ± tightly clustered at stem tips, closely subtended and often overtopped by crowded distal leaves; distal leaves ± thin, usually fringed with numerous weak spines, often ± unpigmented proximally or tinged pink to purplish
var. scariosum
9. Heads usually in ± leafy, racemiform arrays, usually subtended by ± reduced, bractlike distal leaves; distal leaves firm, strongly spiny, usually green throughout
var. coloradense
1. Larger heads 20–35 mm diam.; outer phyllaries elongate, often nearly as long as inner; glutinous ridge narrow, weakly developed
var. cymosum
1. Larger heads 15–25 mm diam.; outer phyllaries usually much shorter than inner phyllaries; glutinous ridge prominent, well developed, appearing dark brown on dry specimens
var. canovirens
Source FNA vol. 19, p. 1. FNA vol. 19, p. 136.
Parent taxa Asteraceae > tribe Cardueae > Cirsium Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Subordinate taxa
C. scariosum var. americanum, C. scariosum var. citrinum, C. scariosum var. coloradense, C. scariosum var. congdonii, C. scariosum var. robustum, C. scariosum var. scariosum, C. scariosum var. toiyabense
C. cymosum var. canovirens, C. cymosum var. cymosum
Synonyms C. hookerianum var. scariosum Carduus cymosus, C. botrys, C. triacanthum
Name authority Nuttall: Trans. Amer. Philos. Soc., n. s. 7: 420. (1841) (Greene) J. T. Howell: Amer. Midl. Naturalist 30: 37. (1943)
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