Flora of North America species comparison

1–several, spreading to erect, bushlike or forming low rounded mounds, gray-tomentose with appressed feltlike trichomes;

branches 0–several, inclined to ascending, stiff.

single, erect, ± glabrous to variably tomentose with coarse, jointed, multicellular trichomes and/or fine smooth trichomes;

branches 0–few, ascending.

blades elliptic to ovate, 10–25 cm, strongly undulate, usually broadly pinnatifid, lobes entire or coarsely few-toothed or -lobed, main spines abrupt, 1–4 mm, faces gray-white-tomentose with appressed feltlike, non-septate trichomes;

basal present or withered at flowering, winged-petiolate;

principal cauline well distributed, winged-petiolate to sessile, gradually reduced, bases clasping with expanded auricles;

distal reduced, spines to 8 mm.

blades broadly elliptic to obovate, 15–40 × 7–15 cm, subentire to coarsely pinnatifid 1/2–2/3 length to midveins, lobes few, lanceolate to triangular-ovate, shallowly lobed or dentate, main spines bristlelike, fine, innocuous, 3–6 mm, abaxial glabrous to villous with septate trichomes or thinly tomentose with jointed trichomes, adaxial faces glabrous or loosely tomentose along midveins;

basal usually absent at flowering, winged-petiolate, ciliate with fine, flexible spines to 8 mm;

principal cauline well distributed, little reduced, bases broadly tapered to clasping, short-decurrent;

distalmost moderately reduced.

0–7 cm.

0–1 cm.

hemispheric or campanulate, 3–4 × 4–6 cm, densely arachnoid.

hemispheric to broadly campanulate, 1.5–2 × 2–3.5 cm, ± densely arachnoid.

tips 3–4 mm.

white to pale yellow, 20–34 mm, tubes 11–15 mm, throats 5–8 mm, lobes 5–8 mm;

style tips 3–4 mm.

pink to purple, 16–17 mm, tubes 8–9 mm, throats 3–4 mm, lobes 4–5 mm.

in 8–10 series, imbricate, linear, abaxial faces without glutinous ridge;

outer and mid bases short- appressed, margins spiny-ciliate, apices long, spreading to erect, spines straight, 2–5 mm;

apices of inner flattened or spine-tipped, serrate to scabrid, sometimes pectinately fringed.

in 5–7 series, subequal, green or tinged purple, linear or linear-lanceolate, abaxial faces without glutinous ridge, outer and middle erect or outer spreading, entire, apices long-acuminate, spines 0–2 mm;

apices of inner phyllaries straight or flexuous, flat.

1–many, erect, terminal on branches in subcapitate to congested, corymbiform arrays, closely subtended by clustered, ± leafy bracts.

1–few, in spiciform or subcapitate arrays.

light–brown to black, 5–7 mm, apical collars whitish, 0.2–0.3 mm;

pappi 15–25 mm.

brown, 4 mm, apical collars not well differentiated;

pappi 12–15 mm.

= 34.

= 68 (Japan).