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few-leaf thistle, mountain thistle, Pacific fringe thistle, remote-leaf thistle, weak thistle

Sacramento Mountain thistle, Sacramento Mountains thistle

Habit Perennials, 20–150 cm, monocarpic; taprooted or polycarpic, perennating by runner roots. Perennials, 100–200 cm; creeping roots.
Stems

usually 1, erect, finely arachnoid-tomentose, sometimes villous with septate trichomes below nodes;

branches 0–10+, slender, usually arising in distal 1/2, ascending.

1, erect, (dark maroon), glabrous;

branches several to many, stiffly ascending, distally nodding.

Leaves

blades linear-oblong to oblanceolate or elliptic, 7–30 × 1–15 cm, unlobed and spinulose to dentate or shallowly to deeply pinnatifid, lobes well separated, linear to triangular-ovate, dentate to deeply lobed, main spines 2–5 mm, slender, abaxial faces green to gray, thinly to densely arachnoid-tomentose, sometimes glabrate, sometimes villous with septate trichomes along veins, adaxial green, glabrous;

basal sometimes present at flowering, sessile or winged-petiolate;

principal cauline mostly in proximal 1/2, winged-petiolate or sessile, bases narrowed, sometimes auriculate;

distal well separated, progressively reduced, becoming bractlike, often unlobed or less deeply divided than the proximal, sometimes spinier than proximal, bases often distally expanded and auriculate-clasping.

blades elliptic, 10–50 × 5–20 cm, 1–2 times pinnately lobed or divided, lobes lanceolate to ovate, main spines slender to stout, 3–10 mm, faces glossy green, glabrous;

basal present at flowering, petiolate or winged-petiolate, divided nearly to midveins;

proximal cauline winged-petiolate;

mid and distal sessile, progressively reduced, less deeply lobed, bases auriculate-clasping;

distalmost linear or lanceolate, bractlike, very spiny.

Peduncles

(0–)2–15 cm.

0.5–15+ cm.

Involucres

ovoid to hemispheric or campanulate, 1.5–2.5 × 1.5–3.5 cm, glabrous to arachnoid-floccose.

(excluding spreading tips) broadly ovoid to hemispheric or campanulate, 2–3 × 2–3 cm, glabrous.

Corollas

creamy white to purple, 18–28 mm, tubes 7–12 mm, throats 5–12 mm, lobes 3.5–7 mm, style tips 4–6 mm.

rich rose-purple, 20–26 mm, tubes 4–5 mm, throats 7–10 mm, lobes 10–11 mm;

style tips 2–2.5 mm.

Phyllaries

in 6–8 series, subequal to strongly imbricate, green, linear to obovate (outer) to linear (inner), abaxial faces with inconspicuous glutinous ridge;

outer and middle bases appressed, margins entire to spinulose-dentate or broad, scarious, lacerate-dentate, spines absent or ascending to spreading, 1–2 mm;

apices of inner sometimes flexuous or reflexed, narrow, flat, entire or expanded, scarious, and lacerate-dentate.

in 8–10 series, strongly imbricate, (dark maroon, drying dark brown or blackish), ovate or lanceolate (outer) to linear-lanceolate (inner), abaxial faces with poorly developed glutinous ridge;

outer and mid bases appressed, margins entire, apices spreading to reflexed, elongate, ovate to lanceolate, 5–20 mm, flattened, ciliolate, adaxially minutely villous with septate trichomes, spines slender, 1–3 mm;

apices of inner flexuous, entire.

Heads

few–many, borne singly or in openly branched in corymbiform, racemiform, or paniculiform arrays on main stem and branches, sometimes also in distal axils, not closely subtended by clustered leaf bracts.

many, nodding, borne 1–few at branch tips, collectively forming open, paniculiform arrays.

Cypselae

tan to dark brown, 4.5–5.5 mm, apical collars differentiated or not;

pappi 13–23 mm.

brown, ca. 5 mm, apical collars not differentiated;

pappi (brown), 18–20 mm.

2n

= 32.

Cirsium remotifolium

Cirsium vinaceum

Phenology Flowering spring–summer (May–Sep).
Habitat Wet soil around calcareous springs and seeps, stream banks, montane meadows, coniferous forest margins
Elevation 2300–2900 m (7500–9500 ft)
Distribution
from FNA
CA; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
NM
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Cirsium remotifolium occurs from the Coast Ranges and valleys of the Pacific Northwest to the western slopes of the Cascade and Klamath ranges, south in the California North Coast Ranges to the San Francisco Bay region. It is closely related to the C. clavatum complex of the Rocky Mountains region. Both have a similar growth habit and some forms variably express the character of broadly scarious, lacerate-toothed phyllary margins. Gray, in naming Cnicus carlinoides var. americanus, included as syntypes both California and Colorado specimens. F. Petrak (1917) treated both the West Coast plants and those of the Rocky Mountains as Cirsium subsect. Americana, recognizing C. remotifolium with several infraspecific taxa plus two other species, C. callilepis and C. amblylepis from the West Coast, and four additional species from the Rocky Mountains. A. Cronquist (1955) rejected Petrak’s subspecies, treating C. remotifolium in a restricted sense, limited to plants of Washington and Oregon without dilated phyllary tips, and circumscribed C. centaureae broadly to include the Rocky Mountains and West Coast plants with dilated phyllary tips. Because of the frequent presence of dilated phyllary tips in C. remotifolium in the restricted sense, Cronquist acknowledged the likelihood of past introgression with C. centaureae in the broad sense.

J. T. Howell (1960b) recognized three species: Cirsium remotifolium, C. acanthodontum, and C. callilepis, the latter with four varieties collectively corresponding to the West Coast representatives of C. centaureae (in the sense of Cronquist). Because of the great similarity of the various West Coast plants and their intergradation, I see no value in recognizing two or more species.

The West Coast and Rocky Mountains plants are clearly related, but are separated by the Great Basin region and there is little chance of current genetic interchange. As is often the case with American Cirsium, genetic enrichment from past hybridization with other nearby species within their respective areas has likely been fertile ground for evolutionary diversification. Different species have contributed genes in the Pacific states and in the Rockies. I have chosen to recognize two geographically-based species complexes, each with intergrading races here treated as varieties. I treat the West Coast plants as C. remotifolium and the Rocky Mountains plants as C. clavatum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Cirsium vinaceum is perhaps most closely related to C. rydbergii. Heads of C. vinaceum are actively visited by hummingbirds and by several kinds of bees (K. A. Burks 1994). Hybrids are known between C. vinaceum and C. wrightii.

Cirsium vinaceum is endemic to the Sacramento Mountains, Otero County. It is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Phyllary margins ciliate with tiny spreading to recurved spines
var. rivulare
1. Phyllary margins unappendaged or dilated, scarious, and ± lacerate-toothed
→ 2
2. Phyllaries narrowly oblong or linear, often ± subequal, all or most without scarious-dilated margins
var. remotifolium
2. Phyllaries oblong to obovate, often strongly graduated, most or all with dilated, scarious, erose to lacerate-dentate margins
var. odontolepis
Source FNA vol. 19, p. 129. FNA vol. 19, p. 163.
Parent taxa Asteraceae > tribe Cardueae > Cirsium Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Subordinate taxa
C. remotifolium var. odontolepis, C. remotifolium var. remotifolium, C. remotifolium var. rivulare
Synonyms Carduus remotifolius Carduus vinaceus
Name authority (Hooker) de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 6: 655. (1838) (Wooton & Standley) Wooton & Standley: Contr. U.S. Natl. Herb. 19: 751. (1915)
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