Cirsium remotifolium |
Cirsium andersonii |
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few-leaf thistle, mountain thistle, Pacific fringe thistle, remote-leaf thistle, weak thistle |
Anderson's thistle, rose thistle |
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Habit | Perennials, 20–150 cm, monocarpic; taprooted or polycarpic, perennating by runner roots. | Perennials (but often appearing biennial), (15–)40–70(–100) cm; rootstocks producing erect, taprooted caudices and rosettes. | ||||||||
Stems | usually 1, erect, finely arachnoid-tomentose, sometimes villous with septate trichomes below nodes; branches 0–10+, slender, usually arising in distal 1/2, ascending. |
usually 1, erect, subglabrous to puberulent and/or tomentose; branches 0–several, stiffly ascending. |
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Leaves | blades linear-oblong to oblanceolate or elliptic, 7–30 × 1–15 cm, unlobed and spinulose to dentate or shallowly to deeply pinnatifid, lobes well separated, linear to triangular-ovate, dentate to deeply lobed, main spines 2–5 mm, slender, abaxial faces green to gray, thinly to densely arachnoid-tomentose, sometimes glabrate, sometimes villous with septate trichomes along veins, adaxial green, glabrous; basal sometimes present at flowering, sessile or winged-petiolate; principal cauline mostly in proximal 1/2, winged-petiolate or sessile, bases narrowed, sometimes auriculate; distal well separated, progressively reduced, becoming bractlike, often unlobed or less deeply divided than the proximal, sometimes spinier than proximal, bases often distally expanded and auriculate-clasping. |
blades ± elliptic, 8–35 × 4–8 cm, divided about halfway to midveins, lobes spreading, triangular, coarsely dentate or with a few broad lobes, obtuse to acute, main spines 1–5 mm, abaxial faces green or gray, thinly tomentose, adaxial green and glabrous to sparingly pilose; basal often present at flowering, spiny winged-petiolate; main cauline reduced distally, bases clasping; distal much reduced, linear-oblong, usually less deeply lobed and often spinier than proximal. |
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Peduncles | (0–)2–15 cm. |
0–20 cm. |
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Involucres | ovoid to hemispheric or campanulate, 1.5–2.5 × 1.5–3.5 cm, glabrous to arachnoid-floccose. |
broadly cylindric to narrowly campanulate, 3–5 × 2–4 cm, loosely arachnoid or ± glabrous, finely short-ciliate. |
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Corollas | creamy white to purple, 18–28 mm, tubes 7–12 mm, throats 5–12 mm, lobes 3.5–7 mm, style tips 4–6 mm. |
red to reddish purple, 30–45 mm, tubes 10–20 mm, throats 10–16 mm, lobes 9–11 mm; style tips 3.5–5 mm. |
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Phyllaries | in 6–8 series, subequal to strongly imbricate, green, linear to obovate (outer) to linear (inner), abaxial faces with inconspicuous glutinous ridge; outer and middle bases appressed, margins entire to spinulose-dentate or broad, scarious, lacerate-dentate, spines absent or ascending to spreading, 1–2 mm; apices of inner sometimes flexuous or reflexed, narrow, flat, entire or expanded, scarious, and lacerate-dentate. |
in 6–8 series, imbricate, outer green, inner purple to red, linear-lanceolate (outer) to linear (inner), abaxial faces without glutinous ridge; outer and mid bodies short, appressed, entire or spinulose-ciliate, apices long-spreading to ascending, entire or spinulose-ciliate or rarely with expanded, fringed appendages, spines straight, weak, 1–3 mm; apices of inner red to purple, straight or rarely twisted, long, flat, entire. |
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Heads | few–many, borne singly or in openly branched in corymbiform, racemiform, or paniculiform arrays on main stem and branches, sometimes also in distal axils, not closely subtended by clustered leaf bracts. |
1–6, borne singly or in corymbiform, racemiform, or spiciform arrays. |
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Cypselae | tan to dark brown, 4.5–5.5 mm, apical collars differentiated or not; pappi 13–23 mm. |
brown, 6–7 mm, apical collars narrow; pappi 25–40 mm. |
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2n | = 32. |
= 32, 64. |
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Cirsium remotifolium |
Cirsium andersonii |
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Phenology | Flowering summer (Jul–Sep). | |||||||||
Habitat | Moist to dry soils, openings in montane woodlands, montaine coniferous forests, aspen groves | |||||||||
Elevation | 1100–2900 m (3600–9500 ft) | |||||||||
Distribution |
CA; OR; WA
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CA; ID; NV
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Discussion | Varieties 3 (3 in the flora). Cirsium remotifolium occurs from the Coast Ranges and valleys of the Pacific Northwest to the western slopes of the Cascade and Klamath ranges, south in the California North Coast Ranges to the San Francisco Bay region. It is closely related to the C. clavatum complex of the Rocky Mountains region. Both have a similar growth habit and some forms variably express the character of broadly scarious, lacerate-toothed phyllary margins. Gray, in naming Cnicus carlinoides var. americanus, included as syntypes both California and Colorado specimens. F. Petrak (1917) treated both the West Coast plants and those of the Rocky Mountains as Cirsium subsect. Americana, recognizing C. remotifolium with several infraspecific taxa plus two other species, C. callilepis and C. amblylepis from the West Coast, and four additional species from the Rocky Mountains. A. Cronquist (1955) rejected Petrak’s subspecies, treating C. remotifolium in a restricted sense, limited to plants of Washington and Oregon without dilated phyllary tips, and circumscribed C. centaureae broadly to include the Rocky Mountains and West Coast plants with dilated phyllary tips. Because of the frequent presence of dilated phyllary tips in C. remotifolium in the restricted sense, Cronquist acknowledged the likelihood of past introgression with C. centaureae in the broad sense. J. T. Howell (1960b) recognized three species: Cirsium remotifolium, C. acanthodontum, and C. callilepis, the latter with four varieties collectively corresponding to the West Coast representatives of C. centaureae (in the sense of Cronquist). Because of the great similarity of the various West Coast plants and their intergradation, I see no value in recognizing two or more species. The West Coast and Rocky Mountains plants are clearly related, but are separated by the Great Basin region and there is little chance of current genetic interchange. As is often the case with American Cirsium, genetic enrichment from past hybridization with other nearby species within their respective areas has likely been fertile ground for evolutionary diversification. Different species have contributed genes in the Pacific states and in the Rockies. I have chosen to recognize two geographically-based species complexes, each with intergrading races here treated as varieties. I treat the West Coast plants as C. remotifolium and the Rocky Mountains plants as C. clavatum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Cirsium andersonii grows in the Cascade Range of northern California south through the Sierra Nevada of eastern California and western Nevada. It has been reported from the mountains of southwestern Idaho, but I have not seen specimens from there. Heads of Cirsium andersonii are actively visited by hummingbirds as well as a variety of insects (P. L. Barlow-Irick 2002). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 19, p. 129. | FNA vol. 19, p. 145. | ||||||||
Parent taxa | Asteraceae > tribe Cardueae > Cirsium | Asteraceae > tribe Cardueae > Cirsium | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Carduus remotifolius | Cnicus andersonii | ||||||||
Name authority | (Hooker) de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 6: 655. (1838) | (A. Gray) Petrak: Bot. Tiddsskr. 31: 68. (1911) | ||||||||
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