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Alameda County thistle, brownie or Alameda County thistle, brownie thistle

Wright's Marsh thistle, Wright's thistle

Habit Perennials, subacaulescent and forming compact, rounded mounds, 5–20 cm, or ± erect and to 70(–90) cm; runner roots producing adventitious buds. Biennials or monocarpic perennials, 100–300 cm; taproots short with many slender, fibrous lateral roots.
Stems

1–10+, erect or ascending, glabrous to thinly gray-tomentose, sometimes villous with septate trichomes;

branches 0 or few, ascending.

usually 1, erect, glabrous to ± tomentose;

branches many, usually restricted to distal part of stem, ascending.

Leaves

blades elliptic to obovate, 5–20 × 3–7 cm, strongly undulate, shallowly to deeply pinnatifid with 3–8 pairs of lobes, lobes linear-lanceolate to broadly triangular, (often longer than 2 cm), closely spaced, spreading, spinose-dentate or lobed, main spines slender to stout, 2–15 mm, abaxial faces thinly to densely tomentose, ± villous with septate trichomes along veins, glabrescent or trichomes persistent, adaxial thinly arachnoid-tomentose and soon glabrescent;

basal usually present at flowering, petiolate;

principal cauline petiolate, progressively reduced distally, bases sometimes decurrent as spiny wings to 1 cm;

distal reduced, similar to proximal.

blades oblong to elliptic, 10–60 × 5–20 cm, unlobed and merely spinulose to irregularly dentate or shallowly to deeply pinnatifid, lobes ± broadly triangular, separated by wide sinuses, obtuse to acute, sometimes coarsely toothed or lobed, main spines slender, 1–3 mm, faces thinly arachnoid, soon glabrescent;

basal often present at flowering, petioles slender, ± winged;

cauline progressively reduced, proximal petiolate, mid and distal sessile, long-decurrent;

distalmost linear to narrowly elliptic, bractlike, spinulose to irregularly dentate or shallowly lobed.

Peduncles

0–10 cm, leafy-bracted.

slender, 1–15 cm.

Involucres

ovoid to hemispheric or broadly campanulate, 2.5–5 (in first-formed heads, often smaller in later heads) × 2.5–6 cm, loosely arachnoid on phyllary margins or glabrate.

ovoid to hemispheric, 1–2 × 1–2 cm, thinly arachnoid, glabrate.

Corollas

white or pale lavender to purple, 25–35 mm, tubes 10–20 mm, throats 7–10 mm, lobes 5–8 mm;

style tips 2.5–4.5 mm.

white to pink-purple, 19–21 mm, tubes 9–10 mm, throats 4–4.5 mm, lobes 5–7 mm;

style tips 2–3.5 mm.

Phyllaries

in 5–10 series, imbricate, ovate or lanceolate (outer) to linear-lanceolate (inner), margins of outer entire, abaxial faces without glutinous ridge;

outer and mid appressed, spines erect or ascending, (0–)1–2(–10) mm;

apices of mid and inner narrowed and scabrido-denticulate or with expanded, spinuloso-serrate or -dentate tips, spineless or spine-tipped.

in 8–9 series, strongly imbricate, green, lanceolate (outer) to linear (inner), abaxial faces with prominent glutinous ridge;

outer and middle appressed, bodies entire, apices acute, spines spreading, slender, ca. 1 mm;

apices of inner often flexuous, acuminate, flat, scabrid-ciliolate.

Heads

1–few, erect, ± crowded, often closely subtended by distalmost leaves.

many, in openly paniculiform arrays, borne singly at tips of peduncles.

Cypselae

brown, 5–6.5 mm, apical collars colored like body;

pappi 20–40 mm.

brown, ca. 4.5 mm, apical collars stramineous, 0.2 mm;

pappi 15–16 mm.

2n

= 32.

Cirsium quercetorum

Cirsium wrightii

Phenology Flowering spring–summer (Apr–Aug). Flowering summer–fall (Aug–Oct).
Habitat Usually dry sites, coastal bluffs, grasslands, oak woodlands, coastal scrub Springs, seeps, marshes, stream banks, often in alkaline soil
Elevation 0–400 m (0–1300 ft) 1100–2600 m (3600–8500 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; TX; Mexico (Chihuahua, Sonora)
[BONAP county map]
Discussion

Cirsium quercetorum occurs in the north and south Coast ranges of California from Mendocino to San Luis Obispo counties. It overlaps in range and habitat with several other thistle species and has been reported to hybridize with C. andrewsii, C. douglasii, C. occidentale, C. remotifolium var. odontolepis, and C. fontinale var. fontinale (F. Petrak 1917; J. T. Howell 1960b). Considerable variation occurs within the range of C. quercetorum, and two of the variants have been given taxonomic recognition as vars. walkerianum and xerolepis. Additional study over the range of the species is needed to determine whether these or other variants should be recognized formally.

Cirsium quercetorum appears to be related to the polymorphic C. scariosum complex. The perennial habit with runner roots of C. quercetorum consistently distinguishes it from the monocarpic C. scariosum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Wright’s thistle occurs from the mountains of south-central New Mexico eastward to the cienegas of the adjacent southwestern Great Plains. Cirsium wrightii is listed by the state of New Mexico as a species of concern. The one known site in Cochise County, Arizona, is apparently historic.

Hybrids are known between Cirsium wrightii and C. vinaceum in the Sacramento Mountains of New Mexico. I have observed hummingbird visits to the heads of both species, though C. wrightii shows none of the apparent adaptations to hummingbirds (P. L. Barlow-Irick 2002) that are seen in such taxa as C. occidentale var. candidissimum, C. andersonii, and C. arizonicum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 19, p. 160. FNA vol. 19, p. 131.
Parent taxa Asteraceae > tribe Cardueae > Cirsium Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri
Synonyms Cnicus quercetorum, C. quercetorum var. walkerianum, C. quercetorum var. xerolepis, C. walkerianum
Name authority (A. Gray) Jepson: Fl. W. Calif., 507. (1901) A. Gray: Smithsonian Contr. Knowl. 5(6): 101. (1853)
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