Flora of North America species comparison

1–10+, erect or ascending, glabrous to thinly gray-tomentose, sometimes villous with septate trichomes;

branches 0 or few, ascending.

usually single, erect, tomentose to ± glabrate;

branches 0–many, usually restricted to distal part, ascending.

blades elliptic to obovate, 5–20 × 3–7 cm, strongly undulate, shallowly to deeply pinnatifid with 3–8 pairs of lobes, lobes linear-lanceolate to broadly triangular, (often longer than 2 cm), closely spaced, spreading, spinose-dentate or lobed, main spines slender to stout, 2–15 mm, abaxial faces thinly to densely tomentose, ± villous with septate trichomes along veins, glabrescent or trichomes persistent, adaxial thinly arachnoid-tomentose and soon glabrescent;

basal usually present at flowering, petiolate;

principal cauline petiolate, progressively reduced distally, bases sometimes decurrent as spiny wings to 1 cm;

distal reduced, similar to proximal.

blades oblong to elliptic, 7–30 × 2–12 cm, unlobed and merely spinulose to irregularly dentate or shallowly to deeply pinnatifid, lobes ± triangular, separated by narrow to wide sinuses, sometimes coarsely dentate or lobed proximally, obtuse to acute, main spines slender to stout, 1–5 mm, abaxial faces arachnoid tomentose, adaxial glabrous or thinly arachnoid;

basal often absent at flowering, petioles slender, ± winged;

cauline progressively reduced, proximal petiolate, mid and distal broadly sessile, bases ± auriculate-clasping or decurrent 1–3 cm;

distalmost linear to lanceolate, bractlike, irregularly dentate or shallowly lobed.

0–10 cm, leafy-bracted.

slender, 3–30 cm (not overtopped by crowded distal leaves).

ovoid to hemispheric or broadly campanulate, 2.5–5 (in first-formed heads, often smaller in later heads) × 2.5–6 cm, loosely arachnoid on phyllary margins or glabrate.

ovoid to hemispheric, 1.5–2 × 1.5–2 cm, thinly arachnoid, glabrate.

white or pale lavender to purple, 25–35 mm, tubes 10–20 mm, throats 7–10 mm, lobes 5–8 mm;

style tips 2.5–4.5 mm.

white to pink-purple, 20–25 mm, tubes 7–10 mm, throats 6–8 mm (noticeably wider than tubes), lobes 4–7 mm;

style tips 3–4 mm.

in 5–10 series, imbricate, ovate or lanceolate (outer) to linear-lanceolate (inner), margins of outer entire, abaxial faces without glutinous ridge;

outer and mid appressed, spines erect or ascending, (0–)1–2(–10) mm;

apices of mid and inner narrowed and scabrido-denticulate or with expanded, spinuloso-serrate or -dentate tips, spineless or spine-tipped.

in 8–10 series, imbricate, green, lanceolate (outer) to linear (inner), abaxial faces with prominent glutinous ridge;

outer and middle appressed, bodies entire, acute, spines spreading, slender, 1–5 mm;

apices of inner often flexuous, flat, scabrid-ciliolate, acuminate.

1–few, erect, ± crowded, often closely subtended by distalmost leaves.

1–many, in openly paniculiform arrays.

brown, 5–6.5 mm, apical collars colored like body;

pappi 20–40 mm.

brown, 3–5 mm, apical collars not differentiated;

pappi 15–16 mm.

= 32.

= 22, 23, 24.