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Wyoming thistle

Beaumont thistle, yellowspine thistle

Habit Perennials polycarpic, 15–60(–90) cm; deep-seated woody tap-roots and caudices. Perennials, 30–90 cm; crown sprouts or runner roots producing adventitious buds.
Stems

1–few, erect or ascending, arachnoid-tomentose or ± glabrate;

branches 0–5+, usually in distal 1/2, ascending.

1–20+, erect or ascending, densely gray-tomentose with non-septate trichomes;

branches 0 or few, usually in distal 1/2, ascending.

Leaves

blades linear to oblong, oblanceolate, or elliptic, 5–25 × 0.6–7 cm, unlobed and merely spinulose or spiny-dentate to regularly pinnatifid, lobes 5–8(–many) pairs, well separated, usually with broad, U-shaped sinuses to crowded, linear to triangular-ovate, ascending-spreading to retrorse, merely spinulose to coarsely dentate or few lobed, main spines 2–7 mm, ± slender, abaxial faces gray to white, usually densely arachnoid-tomentose, sometimes ± glabrate, sometimes villous with septate trichomes along veins, adaxial green, glabrous or less commonly thinly to densely gray-tomentose;

basal often present at flowering, spiny winged-petiolate;

principal cauline well distributed, gradually reduced distally, proximal usually winged-petiolate, mid and distal sessile, bases decurrent as spiny wings 1.5–3.5 cm;

distalmost reduced, ± bractlike.

blades oblong to narrowly elliptic, 10–30 × 2–8 cm, strongly undulate, margins coarsely dentate or shallowly to deeply pinnatifid with 8–15 pairs of lobes 0.5–2 cm, often revolute, lobes ± triangular, closely spaced, spreading, spinose-dentate and cleft into 2–5 spine-tipped divisions, main spines 5–20 mm, yellowish, abaxial faces densely white-tomentose, adaxial thinly gray-tomentose;

basal usually present at flowering, winged-petiolate;

principal cauline sessile, progressively reduced distally, bases ± auriculate to long-decurrent as spiny wings;

distal cauline usually much reduced, less lobed.

Peduncles

0–15 cm.

0–4 cm.

Involucres

ovoid to campanulate, 1.8–2.7 × 1–2 cm, thinly arachnoid-tomentose or glabrate.

ovoid to hemispheric or broadly campanulate, 2.5–4.5 × 2.5–4.5 cm in first-formed heads, often smaller in later ones, loosely arachnoid on phyllary margins or glabrate.

Corollas

pink to purple (creamy white), 18–25 mm, tubes 7–9 mm, throats 5.5–7.5 mm, lobes 4–8 mm;

style tips 3–5.5 mm.

white or pale lavender to purple, pink, or red, 25–45 mm, tubes 8–25 mm, throats 6–17 mm, lobes 6–15 mm;

style tips 2–8 mm.

Phyllaries

in 6–7 series, ± imbricate, green or with dark subapical patch or appendage, linear to linear-lanceolate, margins entire, abaxial faces with narrow glutinous ridge;

outer and middle bases appressed, apical appendages spreading to stiffly ascending, linear-lanceolate to acicular, entire, spines spreading or ascending, stout, 2–7 mm, often flattened;

apices of inner stiffly erect or sometimes flexuous, narrow, flat.

in 5–10 series, imbricate, ovate (outer) to linear-lanceolate (inner), margins entire, abaxial faces with narrow glutinous ridge;

outer and middle appressed, spines spreading, 3–12 mm;

apices of inner often flexuous, expanded and flat, scabrid-margined, sometimes erose, spineless.

Heads

1–few, borne singly or in 2–3-headed clusters in ± congested flat-topped or racemiform arrays at tips of main stem and branches, sometimes also in distal axils.

1–few, in leafy, ± corymbiform arrays.

Cypselae

tan to dark brown, 5–6 mm, apical collars yellow, narrow;

pappi 14–16 mm.

light brown, sometimes with lighter or darker streaks, 6–9 mm, apical collars colored like the body, narrow;

pappi (white or tawny), 20–40 mm, usually noticeably shorter than corolla.

2n

= 34.

= 30, 31, 32, 34.

Cirsium pulcherrimum

Cirsium ochrocentrum

Distribution
from FNA
CO; ID; MT; NE; UT; WY
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; KS; NE; NM; OK; SD; TX; UT; WY; n Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

Cirsium pulcherrimum is closely related to C. clavatum. In southeastern Wyoming and northern Colorado some plants combine foliage and involucral characters of C. pulcherrimum var. pulcherrimum and C. clavatum var. americanum. The inheritance of these characters needs to be examined at the population level to determine whether the intermediates are hybrids or the products of past introgression or incomplete differentiation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Distal leaf face gray- to white-tomentose; cypselae without stramineous apical collar
var. aridum
1. Distal leaf face usually green, glabrous or ± glabrate, but sometimes persistently tomentose; cypselae often with narrow stramineous apical collar
var. pulcherrimum
1. Stems densely leafy, nodes crowded; leaves often long- decurrent; corollas white or pale lavender to purple
var. ochrocentrum
1. Stems leafy, nodes usually well separated; distal cauline leaves clasping, or if decurrent spiny wing usually less than 1 cm; corollas red, pink, or reddish purple (rarely white)
var. martinii
Source FNA vol. 19, p. 125. FNA vol. 19, p. 123.
Parent taxa Asteraceae > tribe Cardueae > Cirsium Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Subordinate taxa
C. pulcherrimum var. aridum, C. pulcherrimum var. pulcherrimum
C. ochrocentrum var. martinii, C. ochrocentrum var. ochrocentrum
Synonyms Carduus pulcherrimus
Name authority (Rydberg) K. Schumann: Just’s Bot. Jahresber. 29(1): 566. (1903) A. Gray: Mem. Amer. Acad. Arts, n. s. 4: 110. (1849)
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